1919] Kofoid-Swcztj: Triclioinini plia campanula 81 



SUMIMARY 



1. This organism has a highlj' specialized flagellate type of 

 structure with a highly developed neuromotor system, the centre 

 blepharoplast connected by a complex system of oblique fibers with 

 the numerous flagella which cover two-thirds of the surface of the 

 body. Besides these fibers the ectoplasm contains an alveolar layer 

 and one of transverse myonemes. Immediately below it in the endo- 

 plasm are the longitudinal myonemes. 



2. The nucleus is submedian in position and part of its chromatin 

 contents is permanently separated as a " heterochromosome " con- 

 tained within a small vesicle. 



3. Nutrition is holozoic but its method of feeding is unknown. No 

 cytostome is present. The endoplasm is divided into two regions, 

 anterior and posterior. The latter region, which is covered only by a 

 thin pellicle and not the thick ectoplasm of the remainder of the body, 

 is usually filled with food particles. These are entirely absent from 

 the anterior region of endoplasm. 



4. The body rounds up at the time of division and the centro- 

 blepharoplast divides, forming a paradesmose, the entire ectoplasm 

 splitting into two parts with it. These act as the centrosome in the 

 succeeding mitotic figures, the spindle fibers arising from the ends 

 of the paradesmose or the centroblepharoplasts. 



5. Precocious splitting of the chromosomes takes place previous to 

 the prophase, forming fifty-two, V-shaped threads. In a pseudo- 

 telosynapsis this number is reduced to twenty-six. These part on the 

 spindle along the line of the original split. 



6. Division of the chromosomes as well as the body is funda- 

 mentally longitudinal. 



7. Trichanynipha campanula is fundamentally flagellate in its 

 morphology and method of division and is in nowise related to the 

 ciliates. It shows a high degree of specialization and development 

 of its neuromotor system which is the most complex one thus far 

 described among the Protozoa. 



8. It is a member of the family Trichonymphidae Kent in the 

 order Hypermastigina Grassi. This order stands near the Poly- 

 mastigina, to the members of which Trichonympha is nearly related 

 both morphologically and in its development. 



Zoological Laboratory, University of California, 

 Berkeley, California. 



Transmitted October S5, WIS. 



