1920] Kofoid: Nomenclature of Human Intestinal Flagellates 155 



patriots Kailliet (1889, 1895), Verdun (1907, 1913), Neveu-Lemaire 

 (1908), and Brumpt (1910, 1913), as well as that of Lenckart and his 

 followers. His names fulfill all requirements under the Code of 

 Nomenclature and have a just claim for con.sideration. 



In view of the internal evidence that Moquin-Tandon had before 

 him Davaine's figures and his acknowledgment, "C'est encore a M. 

 Davaine que nous devons sa eonnais-sance " we award priority to 

 Davaine's Cercomonas hominis over C. obliejua Moquin-Tandon, but 

 establish his Cercomonas elavainei on the reviser's privilege of naming 

 one (but not both) of the two specie.s included under Davaine's C. 

 hominis. 



Trichomonas hominis (Davaine) 



As stated above, the human intestinal Trichomonas, the .smaller 

 species described by Davaine as Cercomonas hominis var. B, is the 

 Trichomonas of the human intestine. The grounds upon which this 

 conclusion rests are as follows. 



(1) The statistical frequency in which Giarclia, Chilomastix, and 

 Trichomonas are revealed in any .series of examinations of diarrheic 

 stools, makes it highly probable that Davaine found Trichomonas in 

 the course of his examinations of choleraic stools. 



(2) There is no evidence that he had found Giardia except the 

 phrase (1854, 1860) "un corpu.scle, ou deux, tres petit, nucleaire" in 

 his account of his var. A (Chilomastix davainei). The two nuclei 

 might have been due either to mitosis or to a confusion with Giardia. 

 There is no additional evidence of Giardia whatever. There is nothing 

 in his account of his var. B to suggest Giardia. We therefore eliminate 

 this genus and have only Trichomonas left as a probability. 



(3) Conclusive evidence that Davaine had found Tnchomonas 

 appears in his firet paper (1854) where two characteristics are noted in 

 the phrases "le filament caudal s'insere un peu sur le cote du corps et 

 en ce qu'il off rait, en outre, un mouvement d'ondulation dans tout le 

 contour. ' ' These are the undulating membrane and the postero-lateral 

 departures of the free end of the lateral marginal flagellum. He omits 

 reference to the undulations in his later (1860) treatise. They are 

 the main features upon which identification is made today of the living 

 Trichonomas. They are not together characteristic of any other 

 human intestinal flagellate. Chilomastix has a cytostomal undulating 

 membrane found only with greatest difficulty, and the posterior end 



