NAIADES OF MISSOURI EAT 
includes more under this group, that is, the genera, Fusconaia, 
Amblema, Megalonatas, Rotundaria, Plethobasus and even some 
species of Pleurobema. 
Quadrula pustulosa (Lea)' 
(‘“Warty-Back,” ‘‘Warty Pigtoe,’’ ‘‘ Pimple Back.’’) 
Plex Viilo Byes. are A and iB. 
183i1—Umo pustulosus Lea, Tr: Am. Phil. Soc., IV, p. 76, pl. VII, 
fig. 7. 
1834— Unio nodulosus Say., Am. Conch., VI. 
1898—Quadrula pustulosa Baker, Moll. Chicago, Pt. I., p. 86, pl. 
XXV. 
ANIMAL CHARACTERS. 
NUTRITIVE STRUCTURES:—-Branchial opening with yellowish 
plumed tentacles; anal smooth; sura-anal very closely con- 
nected—even disconnected—by mantle edge; inner lamina of 
inner gills free from visceral mass; color of soft parts dingy white; 
palpi connected for about one-third of their length. 
REPRODUCTIVE STRUCTURES:—All four gills entirely marsu- 
pial, septa crowded, ovisacs narrow; conglutinates white, leaf- 
like, broken; glochidia large, semi-elliptical, spineless, hinge 
line short and evenly curved, measures 0.230 x 0.300mm. 
SHELL CHARACTERS. 
EXTERNAL STRUCTURES:—Shell ovate-quadrate, higher than 
long, post-umbonal ridge almost horizontal, posterior half pustu- 
late to smooth, dorsal ridge with nodulous costae; beaks pro- 
truding anteriorly sculptured concentrically with upcurved ridges 
posteriorly; epidermis rusty brown with somethimes broad green 
banded rays diverging from beaks. 
INTERNAL STRUCTURES:—Cardinals heavy just under beaks; 
interdentum broad, upright; laterals straight, at right angles to 
interdentum; umbonal cavity compressed, deep; nacre white. 
Sex Length Height Diameter Locality 
OES 2s Xe ub On UX a2 in it (Miss. R., La. Grange, Mo.) 
fot 60 x 60 x 38 ” ( ” ” ” ” ” ) 
OTIS Ati SAT OSIS i OV KIER Ae Gare ” Hannibal, sth) 
No juvenile shells have been obtained for descriptions; this 
species is so rare in its typical form for this State that adult shells 
have been secured with difficulty. 
' The name of this species should read Q. bullata (Raf.) if we accept 
Rafinesque’s evident description of it in his Monograph (1820, p. 41). 
