386 Stainforth: The Guests of Yorkshire Ants. 
and on dead flies), and secrete from the abdominal hair-tufts 
a sweet substance, to ants both palatable and stimulating. 
C. testaceus has been taken in many places in the south 
and west of England, as in the Isle of Wight, Somerset, Oxford 
and N. Wales; and in north, in the Tweed and Forth districts. 
Its discovery in the intervening area of Yorkshire could thus 
have been prophesied, and careful search in nests of Donis- 
thorpea spp., will probably prove its occurrence in other districts 
in Yorkshire. I regret that a search during the present year 
in numerous nests of D. flava, in Holderness and the Wolds 
district of the East Riding, has had a negative result, but I 
have not yet given up hope of finding it there. 
Atemeles emarginatus, the remaining true ant-guest recorded 
for Yorkshire, has been found near Doncaster. This is appar- 
ently the northernmost record, as Fowler* says he knows ‘ of 
no locality further north than Lincoln,’ where he took it with 
Formica fusca. It is possible that this species is extending 
its range northward, and if so we may be hearing of its occur- 
rence in other parts of the county. I have not been successful 
in my search for it in the Hull district, which unfortunately, 
is somewhat out of the main stream of migration from the 
south. 
The remaining four British symphiles have a southerly 
distribution, and unless a northward spread of the species takes 
place before the increase of coal mines and the greater extension 
of agricultural and industrial areas raise an insuperable barrier 
to their advance, it is scarcely probable that these will ever 
become Yorkshire species. I have shown on the accompanying 
sketch maps the relative distribution of the six British true 
ant-guest beetles. It should be added that Claviger longicornis 
has only recently been added to the British list on the authority 
of specimens captured near Oxford by Commander Walker in 
1906. 
: We may, perhaps, include among true guests, the species of 
aphides which are the normal inhabitants in the nests of many 
ant species, such as D. flava. I found aphides in abundance in 
the nests of this ant at Weedley, and submitted specimens to 
Prof. F. V. Theobald who kindly informs me that they are the 
ground form of the Elm Gall Aphid Tetraneura ulmi. These 
occurred, clustered round the grass roots which penetrate the 
tunnels of the nest, in company with groups of ants. 
INDIFFERENTLY TOLERATED TENANTS.—Of the ‘ indiffer- 
ently tolerated tenants’ (synoeketes), or, in Donisthorpe’s 
phrase, ‘indifferently treated lodgers,’ several have been 
* Coleoptera of the Brit. Isles, vol. II., 1888. 
+ See ‘The Myrmecophilous Coleoptera of Great Britain,’ by H. St. 
J. K. Donisthorpe, Col. Brit. Isles, vol. VI., 1913, pp. 320-330; and 
Rep. of the Lancs. and Ches. Ent. Soc., for 1905, pp. 35-44. 
Naturalist, 
