118 THE AMERICAN MIDLAND NATURALIST 
CRYPTOPHILA PUDICA var. KNAPKEI var. nov. 
(Illustration 3 of Plate II.) 
2 
Radices usque ad 2 mm. crassae parum coopertae micor- 
rhiza. Scapus subterraneus erectus 1.5-2 cm. altus exclusa 
inflorescentia. Inflorescentia supra terram paululum in terra 
infixa, capitulum; flores 3-9; rachis minus quam 1 cm. longa. 
Bacca circa 5 mm in diametrum. 
The small group of plants known as Monotropaceae, Mono- 
tropeae, Monotropoideae, Hypopityaceae and Hypopithieae 
has been repeatedly changed with regard to rank and position 
in the systems of classification, due rather to minor points, for 
its relation to the large body of the Ericals is well established. 
Even at present no agreement has been reached and thus we 
find the group considered a distinct family, or a subfamily of 
the Pyrolacia, or again a subfamily of the Hricacea according 
to the opinions of different authors. As the group itself 
retains its integrity regardless of supposed rank or position, 
the matter is of no consequence in the present case. 
However, in order to determine the position of the 
new genus within the group of monotropoid genera we are 
concerned, with the subdivision of the group into tribes of 
which two have been recognized hitherto, the Eumonotropeae 
and the Pleuricosporeae. Cryptophila differs.from the Pleuri- 
cosporeae mainly in the ovary structure and the fruit char- 
acter, and from the pleuricosporeae mainly in the anthers, 
in the presence of a manifest disk and in fruit. The arrange- 
ment thus becomes: 
Tribe I. Eumonotropeae: Ovary several-celled; placen- 
tae projecting from a central columnella; fruit a loculicidal 
capsule; disk present or none. It insludes the genera Allo- 
tropa, Pterospora, Sarcodes, Monotropsis, and Monotropa. 
Tribe II. Pleuricosporeae: Anthers opening by longitu- 
dinal slits; ovary one-celled or falsely several-celled through 
fusion of the 2-lamellate placentae; placentae parietal; fruit 
a loculical capsule; disk none. It contains the genera Pleuri- 
cospored,. N ewberrya. and. Cheilotheca. 
Tribe IIT. Cryptophileaes Arithers opening: se ‘pores; 
