UNIVERSITY OF CALIFORNIA PUBLICATIONS \ “ 
2e P 
ZOOLOGY 
Vol. 17, No. 9, pp. 115-125, 3 text figures February 3, 1917 
NOTES ON THE SYSTEMATIC STATUS OF THE 
TOADS AND FROGS OF CALIFORNIA 
BY 
CHARLES LEWIS CAMP 
(Contribution from the Museum of Vertebrate Zoology of the University of California) 
The large series of amphibians contained in the collection of the 
Museum of Vertebrate Zoology at the University of California has 
been at the writer’s disposal for study and has furnished opportunity 
for looking into the status of certain California toads and frogs. 
Through the courtesy of Dr. Leonhard Stejneger, Head Curator in 
the Department of Biology of the United States National Museum, 
the writer was accorded the privilege of examining specimens in that 
museum, where the types of nearly all of the western species of frogs 
and toads are preserved. 
As a result of this study it is found necessary to consider the two 
long recognized forms, Bufo halophilus and Bufo boreas, as subspecies 
of one species, and to put Bufo columbiensis under the synonymy of 
Bufo boreas boreas. Two new races of that confusing species, Rana 
boylivi, are described, and two other western frogs, Rana draytonw and 
Rana aurora, are placed together as subspecies of aurora because of 
the recognition of intergrades from the coast region of northern Cali- 
fornia. Notes are also given on the occurrence in California of Bu/jo 
boreas nelsoni and the two subspecies of Rana pretiosa. 
Toaps 
Examination of about 370 toads of the several forms closely alhed 
to Bufo boreas, from Alaska, Vancouver Island, eastern Washington, 
western Oregon, northern Nevada, and California, has convinced the 
writer that most of the characters that have been employed by various 
authors, including Baird and Girard (1854a, pp. 174-175, 378-880, 
18546, p. 301), Girard (1858, pp. 74-80, atlas, pl. 5, figs. 4-9, pl. 6, 
