350 Gates. 
at o, as in O. Lamarckiana, and others in class 5, as in typical 
O. rubrinervis, and others at 2 or 3. Hence there is here probably 
no inheritance of color pattern between classes 0 and 5, although a 
knowledge of another generation would be necessary, to be quite 
certain on this point. 
The O. Lamarckiana from DE VRIES’ cultures also frequently shows 
a range of color pattern from o to 5. This color pattern is also 
characteristic of the buds of O. laevifolia, and often occurs in O. brevi- 
stylis and less frequently in O. nanella and O. gigas, but is rarely 
found on O. lata or O. semilata. 
Another series of observations was made in 1909, on the sepals 
of a form having buds and some other characters quite similar to 
O. rubrinervis, but with rather more color developed. The form in 
question is the offspring of a single individual (No. 25) in a culture 
of Oenotheras from near Liverpool, England. 
A total of 111 plants were grown in Igog from this individual, 
and they proved to be a nearly constant race, the chief variation 
displayed being in the production or omission of a rosette stage, 
twenty plants forming no rosette at all. The characters of this race 
make it probable that they are complex hybrids between O. grandiflora 
and one of the O. Lamarckiana forms, possibly O. rubrinervis. The 
bud characters are mostly intermediate between O. rubrinervis and 
O. grandiflora Ait. 
The observations on the color pattern of the buds were carried 
on from July r9™ until August 14", r909 nearly all the buds being 
examined which appeared during this time, which is the height of 
the blooming season. The results for each individual studied, are given 
a separate line, showing the varying distribution in classes in different 
individuals (see table 8). There was no marked change in the mode 
of an individual during the season. The intermediates between classes 
have been removed, half to each adjacent class, as with the O. rubri- 
nervis data. 
In every case there is a single mode when classes 4, 5 and 6, 
which show the same extent of color pattern, are summed. The total 
shows that the plus variations are in this case much more numerous 
than the minus variations, the mode falling in class 7 in several 
individuals, and no buds occurring in classes o—2. In the case of 
the O. rubrinervis buds, the mode for the whole population of buds 
(see table 1) is also in class 4—5—6, but the minus variations are 
more numerous than the plus variations. 
