38 BOTANICAL GAZETTE [yJuLy 
cell, but the latter appears before the first division in the former, 
a two-celled fruiting branch being accompanied by a correspond- 
ing vegetative one. When, in rare cases, an older fruiting 
branch was found alone and apparently terminal, as in fig. 3, it 
was not always possible to determine whether it was an abnormal 
state, or whether the sterile cell had been severed by accident. : 
The mother cell of the fruiting branch is hemispherical, and — 
is richer in protoplasm than the joint from which it arises. It is 
soon divided by a wall parallel to its base into a flattened, more 
or less irregular, hemispherical upper cell and a lower irregularly 
oblong cell. 
A cell is now cut off from the upper cell by a wall oblique 
to the axis of the branch; as is seen in jig. 3. When viewed in 
some positions this wall appears parallel to the first, as is shown _ 
in fig. 4. Careful focusing in this case, however, showed that — 
the third cell lies partially under its mother cell and is separated : | 
from it by an oblique wall. This may be the terminal cell of 
Nageli and Janczewski. It lies in this particular case, however, 
on the side of the branch towards the vegetative cell, while 
Janczewski, judging from his figures, though he does not explain 
his view point, terms cells in this position lateral, and these, 
according to him, arise after the terminal and the anterior cells a 
in G. corallina., a 
Spalding $ calls attention to the “ irregularity in both the num- 
ber and position of peripheral cells in G. Bornetiana.’’ The — 
present study has confirmed his conclusion, and hence has made 
the use of such terms as lateral and anterior appear of doubtful 
Propriety in this species, at least. 7 . 
Fig. 5 represents the same stage as fig. 4, but with the 
vegetative cell at one side. Here the first two peripheral cells 
are plainly “lateral” according to Janczewski’s use of the term, 
but this was not the case in all the preparations studied. 
Before the appearance of a third peripheral cell, one of the , 
two first cut off usually divides into two cells, of which the — 
5 Development of the sporocarp of Griffithsia Bornetiana. Abstract; Proc. A. A. 
A. S. 39: 327. 1890. 
