Se eT eee TN ee Te 
Se Bia: ha 
ee See ee pee 
eee a! RI ree Re en ney Th eee 
1896] CYSTOCARP OF GRIFFITHSIA 39 
upper is the smaller (fig. 6). A third peripheral cell is now 
cut off, after which a second peripheral cell frequently divides 
into two cells, making at this time two two-celled branches and 
one of one cell arising from the centralcell. The term central cell 
in this paper, as in the one quoted,’ refers to the second cell of 
the fruiting branch, from which the peripheral cells are cut off, 
not to the cell giving rise to the spores, to which the term has 
been applied by Schmitz? and Hauptfleisch,® but which is here 
designated as the placental cell, a name used in this sense by 
Janczewski. This seems to be the normal order, although cases 
were frequently observed in which only one of the branches 
consisted of two cells, while in some cases no “appendicular”’ 
cell was found in the entire branch. Dr. Farlow does not refer 
to this division of part of the peripheral cells, and whether it is 
not a constant feature in this species, as Janczewski holds it to 
be in G. corallina, or whether the appendicular cell, being spe- 
cially liable to be severed in manipulation, is thus frequently lost 
sight of, is not easily determined. 
The fourth peripheral cell, to which both Farlow and Janc- 
zewski refer, was in no case observed in the earlier stages, and 
its presence was not demonstrated in later ones. This coincides 
with Spalding’s conclusion in regard to the variability in the 
number of these cells. 
The peripheral cells usually present a flattened hemispher- 
ical form in optical section, with beak-like projections of proto- 
plasm where they join the mother cell. When lying over the 
mother cell, the outline is broadly elliptical. They remain 
smaller than the central cell, which can readily be distinguished 
until nearly the last stages of the developmental process by its 
size and its tetrahedral form, produced by the oblique walls 
6 Farlow, l. c. 
7 Untersuchungen iiber die Befruchtung der Florideen ; Sitzb. d. konig. pr. Akad. d. 
Wiss., Berlin,—: 215. 1883. English translation of same by W. S. Dallas; Ann, and 
Mag. Nat. Hist. 13:1. 1884. 
® Die Fruchtentwickelung der Gattungen Chylocladia, Champia, und Lomentaria; 
Flora 75: 307. 1892. 
