278 BOTANICAL GAZETTE [ OCTOBER 
tem of tissues, although pointing out that its hyphz are coarser, 
more highly refractive, more loosely arranged, and more irregu- 
larly intertwined than is the case in the rest of the medullary 
system. In all of these points of difference which have been 
enumerated, the tissue in question agrees with that forming the 
surface of the egg. As it is also more intimately connected 
with such cortical tissue than with the medullary tissue, stains 
the same as the former, and in the youngest stages (figs. 7 and 
2) is the direct continuation upward in the egg of the cortical 
tissue of the mycelial strand, it should be regarded as belonging 
to the cortical system. It will be referred to in this article as the 
tissue of the cortical plates, as in my earlier paper on Anthurus. 
In the earliest stages the cortical layer is clusely adnate 
to the medullary layer. In the stage of fig. ¢ separation of 
these two layers begins along the inner edge of each cortical 
plate, the two tissues seeming to be pulled slightly apart, 
although very numerous hyphal connections still exist between 
the two surfaces. In the older stage of jig. 5 this sepate 
tion has become more complete and a decided fissure has 
‘ been produced between the two systems along the inner edge # 
the cortical plate (C’). While it is possible that the rapid 
growth of the medullary lobes (G) and their distention with the 
gelatinous accumulation may have carried the cortical system 
outward bodily and, to some extent, may have loosened the 
cortical plates from their connections along their inner edges, 
it seems more probable that the separation has been callst 
chiefly by changes in the medullary structure facing against the 
cortical plates, as shown in fig. 5. Medullary hyphe reaching 
to the edge of a cortical plate become swollen at the outer end 
and become arranged side by side in a palisade-layer. i 
of a similar nature crowd their way in between the members © 
this palisade-layer and so increase its surface that the layet 
becomes thrown into folds (¢, fig. 5) and torn away shine : cf 
nections with th ): 10 
e edges of the plates (C’) s kind 
sections of this stage occasional hyphal connections of thi 
still persisted, 
