282 BOTANICAL GAZETTE [OCTOBER 
into the cavity which has formed there, and spreads about in it, 
and between the medullary masses (4),and over the surfaces of 
those masses and on that of the cavity. For some reason (which 
may, perhaps, be proximity to a supply of available food) these 
cortical hyphe find the conditions for their further differentiation 
most favorable on the surfaces of the masses of gelatinous tissue, 
and they become closely adherent to such surfaces and grow 
very luxuriantly there. Still, many of the hyphe are found in 
the more open space, running irregularly through such spaces or 
crossing from one side to the other. It is this tissue which 
finally fills the spaces between and ‘about the gelatinous masses 
and becomes the chamber walls of the receptaculum. 
An older stage of the rudiment of a column of the recep- 
taculum and of the tissues about it is shown in cross section in 
fig.7. On the right is the tissue of the gleba with its chambers 
lined by the hymenial layer (7). On the left is the inner 
edge of a cortical plate (C”);-its hyphae may be seen passing 
among and against the medullary masses (4) of the future 
chambers. These cortical hyphe are becoming laterally inflated 
in this region and are plainly recognizable as early stages of 
pseudoparenchyma. Along the surfaces of the larger cavities, 
the development of these hyphae is giving rise to pseudoparer 
chymatous plates (f,f) which in later development have the 
intervening space more filled in with this tissue, forming 2 MOF 
compact partition wall. Near the gleba, large masses (0) of the 
tissue of the chambers are connected, as in fig. 6, with the tramal 
tissue (7) and with the tissue on the inner flanks of the column of 
the receptaculum, tissue just on the border between the gleba - 
the gelatinous layerof the volva. At places in the cross-section 
where the two systems of tissues come into most intimate CoM 
tact and where the plane of the section may happen to gr 
cross-section of the pseudoparenchymatous hyphe, it becomes 
very difficult to determine the true relations to each other of the 
. Te € 
adnate tissues # and 6. I have examined such places with re 
utmost care and find no connections between the tissues of oe 
aes and: the pseudoparenchyma of the walls. 1 
