286 BOTANICAL GAZETTE [ocTORER 
becomes gelatinous and finally disappears, giving rise to the 
chamber cavities. 
5. That since the tissue of the knots is found to be in con- 
nection with the tramal tissue of the gleba at some points, there- 
fore it follows that the receptaculum and the gleba are homolo- 
gous, that the tissue of the chambers of the former is identical 
with the tramal tissue of the latter, and that the pseudoparen- 
chymatous walls of the receptaculum chambers are homologous 
with the hymenial layer of the gleba. 
Comparison of Fischer's figs. 5 and 7 with my figs. 6 and7 
shows that we both see the same general structure and that our 
difference is in its interpretation. That Fischer’s interpretation 
of the structure is not the correct one seems to me to be shown 
by the following considerations: 
1. The existence in the Clathree of the supposed hyphal — 
knots is very doubtful. My experience is that if the attention 
is fixed upon one of them, and this is then followed through 
section after section of the series, the supposed ‘‘knot’’ is soon 
found to be connected with other masses, and so on with the 
main medullary mass. 
2. The tissue of the cortical plates C’ (Zwischengeflecht of 
Fischer) does not form hyphal knots. About small medul- 
lary branches located in the densest portion of the tissue Cc. a 
at 2, fig. 9, it often happens that the plane of the section will 
be unfavorable for displaying the true relations of the two closely 
adnate tissues. At such a place one needs all of the aid to be 
had from good double-stained preparations. 
3. If pseudoparenchyma of the chamber walls arises from 
the hyphe of the chambers by the projection of the swollen 
hyphal tips outward into the narrow intervening spaces, it ought 
to be found distinctly connected with the tissue of the chambers, 
as in the Phallez, and its short hyphez should also be found 
projecting from the sides of the chamber masses into the neh 
ToW spaces in directions perpendicular to the surfaces of the 
chambers. Such a direction of the pseudoparenchymato® 
hyphz should be shown distinctly, as it is in the Phallez. - 
