Se eT ee ea ee eae eee 
1896] THE PHALLOIDE OF THE UNITED STATES 287 
some parts of the section, where the pseudoparenchymatous 
hyphe are closely crowded against the chamber tissue, and 
where the plane of the section has cut these hyphe transversely, 
one must be on his guard against mistaking the cut ends of 
adjacent pseudoparenchymatous hyphe for pseudoparenchyma- 
tous bodies standing out perpendicularly from the sides of the 
chambers. Such closely crowded cut ends, with their looser 
arrangement toward the middle of the space, form a block-like 
structure very well adapted for giving the familiar optical illu- 
sion of radiation. 
I fail to find hyphal connection between the pseudoparen- 
chyma and the tissue of the chambers, nor is there an arrange- 
ment of the pseudoparenchymatous hyphe perpendicular to the 
surface of the chambers. On the contrary, wherever the plane 
of the section discloses the arrangement, these hyphz are found 
lying parallel with the surfaces of the chamber masses, as shown 
In figs. 8 and 9g. If Fischer’s theory were true, such an arrange- 
ment should not be found at any point. 
4. Since the homology claimed between the pseudoparen- 
chyma and the hymenial layer of Clathrus depends upon the 
origin of pseudoparenchyma from the tissue of the chambers, it 
ceases to be tenable. 
The theory of the origin of the receptaculum enunciated by 
Ed. Fischer has been accepted by A. Miller? in his interesting 
a instructive work on the Brazilian Phalloidew. His investi- 
Sation seems to have been completed and his work in process of 
publication when my paper on Anthurus reached him, as his 
references to it are in footnotes. 
SUMMARY FOR THE CLATHRE&. 
lar ane earliest stage, the egg consists of cortical and medul- 
7. stems continued upward from the mycelial strand. 
the ES cottical layer gives rise to the outer layer of the reyes | 
ulum ag plates, and the pseudoparenchyma of the receptac 
ee medullary portion gives rise to the gelatinous masses 
* Brasilische Pilzblumen. Jena, 1895. 
