288 BOTANICAL GAZETTE [OCTOBER 
of the gelatinous layer of the volva, to the gleba, and to the 
gelatinous tissue of the chambers of the receptaculum. 
In such differentiation, the gelatinous masses of the volva 
are the first to be set off. They arise by growth outward into 
the cortical region of the hyphe from several areas of the 
meduilary mass. : 
The masses of cortical tissue which separate the medullary 
gelatinous outgrowths from each other retain their connection 
with the outermost layer through all the later development of 
the egg. They may be called cortical plates; their position is 
indicated by shallow furrows on the surface of the egg. 
Along the inner edge of each cortical plate, a cavity forms 
by the separation of the two systems of tissue in that region. 
The medullary tissue facing these cavities gives rise to the 
gleba and to branching hyphal masses, which project outward 
and upward into the cavities and become the chamber tissue of 
the receptaculum. 
The receptaculum is formed through the joint participation of 
the two systems of tissue and differs in its origin and mode of 
formation from that of the Phallee. Its chambers have the 
position originally occupied by a branched structure of medullary 
origin, while its pseudoparenchymatous walls form by ingrowth 
of cortical tissue about and between the chamber masses. 
The pseudoparenchyma of the walls is not homologous with 
the hymenial layer. 
The elongation of the receptaculum in C. columnatus begins 
at the base. After elongation of the receptaculum, the gleba 
hangs suspended from the arch of the receptaculum by medullary 
tissue constituting chamber masses of the receptaculum. 
RELATIONSHIP OF PHALLEA AND CLATHRE#. 
The Phallez and the Clathrez, the two usually accepted - 
orders of the Phalloidea, resemble each other: 
1. In gross structure. Both have a volva of the same 
general structure, which becomes ruptured by the same means 
and from which there issues the receptaculum. 
