290 BOTANICAL GAZETTE [ocroBER 
side of the peripheral portions of the sheaflike head, while in 
the Clathree its portions alternate with the gelatinous lobes of 
the volva. (c) The basidia of the young hymenium of the 
Phallez face towards the axis of the plant; in the Clathree 
they face in the opposite direction, towards the periphery. 
(d@) In the Phallez, tissue of medullary origin gives rise 
both to the pseudoparenchyma of the receptaculum and to the 
tissue of its chambers; in the Clathrez, cortical tissue contin- 
uous with that of the mycelial strand forms the pseudoparen- 
chyma,™ while the tissue of the chambers is of medullary origin 
and connection. 
The marked resemblances that have been pointed out between 
the mature stages of the Clathree and Phallee have been 
regarded in systematic botany as sufficient for joining them into 
a well-marked order, the Phalloidex. The difference in the 
position of the gleba, whether on the outside of the receptacu- 
lum or within it, has been made the basis of their separation as 
suborders. 
Rehsteiner’s work ” on Hysterangium clathroides and Hymeno- 
gaster decorus, both belonging to the H ymenogastree, led him, is 
the ground of the position of the hymenium and the direction 10 
which it faces, to point out a probable origin of the Clathrex 
from a Hysterangium-like ancestor, and a possible origin of the 
Phalleze from one of the Hymenogastree near Hymenogastet, @— 
view Ed. Fischer has also since expressed.*3 ee ee 
Thaxter’s account of Phallogaster saccatus*+ indicates that this 
may be a species somewhat closer to the Clathrea than Ayster 
angium clathroides is, on account of having a gelatinous layer 
cal in 
lial A is all oe Ser a fn 
oe That the pseudoparenchyma of the receptaculum of the Clathre is ae evil 
origin 1s not strange. A. Miller states that in Clathrus columnatus and agent 
Ta: 
rhacodes the cortical layer of the mycelial strands is pseudoparenchymatous (B 
i P 
33°: 44. 1893. 
™4 Note on Phallogaster saccatus. Bot. GAZ. 18:117. pl. g. 1893- 
