vital THE PHALLOIDLZ OF THE UNITED STATES 291 
between its gleba and peridium that is to be regarded, probably, 
as the forerunner of the gelatinous masses in the volva of the 
Clathree. 
A. Miller’s description of his Protrubera Maracuja makes a 
still closer connection. He finds not only gelatinous masses 
next to the peridium, but also finds them arranged alternately 
with reference to the glebal masses, as in the Clathree, and sep- 
arated from each other by cortical plates along whose inner 
edges the glebal masses develop. The development of the cor- 
tical plates from the peridium, and their connection with it, are 
made very probable by his figs. 2-5, pl. V1. 
My studies, showing that the pseudoparenchyma of the 
receptaculum of the Clathrez is of cortical (peridial) origin, sup- 
ply another link in the chain connecting the Clathree with Hys- 
terangium. By making known the early differentiation of the 
Phallez, and by showing that this subfamily differs wholly from 
the Clathrea in the development of the receptaculum, they make 
it certain that the Phallez, although as highly a differentiated 
subfamily, cannot have arisen from the Clathree. 
It seems safe to conclude: 
1. That the Phallez are not directly related to the Clathreez. 
2. That both subfamilies have arisen from lower forms out- 
side their family. 
: 3. That the Phalloidez consist of two parallel series of forms, 
Which, through variations from unlike starting points, have 
— to highly specialized structures adapted to the same 
ends 
Mi 
eIPPLEBURY COLLEGE, MIDDLEBURY, VT. : 
EXPLANATION OF PLATES XI AND XII. 
ots “shebie were drawn with the aid of an Abbé camera lucida. The = 
plate; Phe are common to all the figures: C, cortical layer; C", corticé 
» G, gelatinous layer of volva; GZ, gleba; H, hymenium ; M, medul- 
young stages; 3, tissue of the chambers ; #, inner layer of the 
TP arenchymatous wall of the chambers; 4, tramal tissue. 
'S Ibid, 10, 
