324 BOTANICAL GAZETTE [ocToRER 
ular and usually somewhat vacuolated. As they grow they 
become slightly constricted at the base. Certain ones advance 
more rapidly than others, so that in a single perithecium all | 
stages of development may frequently be seen. The “bouquet” 
of variously sized asci lies embedded in the loose surrounding 
parenchyma, a part of which persists until the asci are quite 
mature. A distinct layer of this tissue is present between the 
bases of the asci and the basal part of the wall. In the center 
of the perithecium, between the asci, these vegetative cells 
become disorganized and probably furnish nourishment for the 
growing asci. 
Meantime the wall has also undergone some changes. With 
the growth of the asci the entire sporocarp enlarges, the cells of 
the outer wall become thicker walled and darker, while those of 
the inner wall undergo still more lateral compression. At a 
point opposite the base the ostiolum begins to appear. Is 
beginning is marked by a slight protuberance, beneath which 
first the cells of the inner then those of the outer wall begin © 
separate schizogenetically. The ends of these elongated innet 
wall cells after separation round off and look like filaments 
verging toward the canal of the ostiolum. As the neck increase 
in length these filaments also lengthen and extend up into the 
neck canal parallel with its sides. The growth of the ostiolum — 
continues after the development within is quite complete: 
Indeed the asci often are broken down, leaving the spores ee i 
within the cavity before the neck reaches its normal length. Se 
The nuclei of the mycelium have already been noticed. 
They seem essentially the same in structure as those wee : 
Teichospora, consisting, so far as could be seen, of a nucleo ce 
ina clear circular court. The fact that no linin has bee? wee 
in the nuclei of the mycelium is probably due to the we 
of these nuclei and the fact that they are seen through me 
of the mycelium. In the ascus the nuclei of Ceratostom@ ; 
smaller than in Teichospora and fewer details were we 
The drawings were reproduced free-hand, as those made ] 
the camera lucida were too small for satisfactory illustration. ~ 
