1896 PROCARP AND CYSTOCARP OF PTILOTA 359 
trichogyne. However, before this takes place the basal cell of 
the branch, that which is attached directly to the terminal seg- 
ment of the procarpic branch (fig. 4,c) gives rise to a cell 
laterally. This condition is shown in fig. 5, cell 10’*. From 
this last is developed another branch of three or four cells. The 
two-celled stage is shown in fig. 7, and the three-celled con- 
dition in fig. 8, in both cases numbered 10’*. By the elongation 
of the terminal cell of this second branch the structure becomes 
a procarp. We have now two procarps of three or four cells 
each, lying side by side, united to the cell that was first cut off 
from the tenth segment. By examining the later stages of this 
pair of procarps (figs. 7, 8, and g) it will be seen that the 
trichogyne of the first procarp (procarp 10’) develops before 
the second. In fact it is usually the first of all the trichogynes 
in the group of procarps to mature. 
It is well to call attention now to the fact, which will receive 
more detailed treatment later in the paper, that the cystocarp of 
P. serrata, in all cases that the writer has examined (some 112 in 
number), has always developed from the cell at the base of the 
pair of procarps. This carpogenous cell (¢ in jigs. 3-9) is the 
first to be derived from the terminal segment of the procarpic 
branch, and therefore next to the basal cell of procarp no. 11 is 
the oldest cell of the group. 
There remain to be considered the two procarps that lie 
between the inner procarp and the pair on the outside. As is 
shown in jig. To, these two procarps are attached laterally to 
the terminal segment of the procarpic branch in such a manner 
that when viewed from the side one appears in front and the other 
behind this cell. Their development is precisely like that of the 
Procarp on the inside of the group. A cell is cut off first on one 
Side of the terminal segment and then on the other side (figs. 5 
and 6, cell 10"). Each of these two cells then develops into a 
Procarp of three cells in exactly the same manner as the other 
Procarps develop. This is well shown in the figures of later 
Stages ( figs. 7,8,and g). These two procarps are the last of 
© group to mature. 
