1896 | PROCARP AND CYSTOCARP OF PTILOTA 361 
inner zone was much more strongly tinted than the outer. Even 
in fresh material and unstained specimens, the inner zone appears 
of a denser consistence. A most interesting peculiarity of this 
cell-wall is a series of radiating strands which arise from the 
edge of the inner zone and pass through the outer zone to the 
outer edge of the cell-wall (figs. 4, 5, 6, etc.). These radiating 
strands stain about as deeply as the inner zone, and appear to be 
of the same substance. This complex cell-wall is very common 
around the developing procarps, and is sometimes found, but not 
insuch a characteristic form, at the apical cell of the pinnae. 
The peculiar swollen appearance of the outer zone suggests the 
phenomena of gelatination, and to test this point the writer 
treated specimens with a hot solution of potassic hydrate. The 
consistence of the outer zone was quite unaffected by this treat- 
ment, instead of swelling or dissolving as substances of a gelat- 
inous nature would have done. The writer was quite unable to 
obtain a cellulose test (using iodine and sulphuric acid), either 
with this curious cell-wall or with the ordinary cell-walls of this 
plant. But there are reasons from its general appearance and 
reaction towards stains for believing it to be at least closely 
related to cellulose, if it be not that substance. 
Adopting the terminology of Bornet and Thuret we 
divide the procarps into three portions: (1) the trichogyne, (2) 
the carpogenous cell, and (3) the portion of the procarp lying 
between these two, consisting of one or two cells, which we may 
call the trichophoric apparatus (/appareil trichophorique ). 
e know certainly of but one carpogenous cell in the group 
of procarps, and this is the basal cell of the first procarp of the 
pair on the outside of the group. However, it is probable that 
the basal cell of each procarp is morphologically a carpoge- 
Nous cell. At all events the following remarks on the structure 
of the cell that does give rise to the cystocarp are equally 1 
oe the basal cells of all procarps. The carpogenous cell at the 
time when the trichogyne is mature is the largest one in the pro- 
carp. It is slightly tinted with the red color of the Floridee, 
but a well defined chromatophore cannot be made out dis- 
may 
