362 BOTANICAL GAZETTE [NOVEMBER 
tinctly. The central portion of the cell is a cavity containing cell 
sap, and the protoplasm with the irregular chromatophore forms 
a layer next the cell-wall. There is a distinct nucleus imbedded 
in the protoplasm, and as a rule a well defined nucleolus is 
apparent in specimens stained with haematoxylin (figs. zz and 12, 
c). The carpogenous cell is connected below with the terminal 
segment of the procarpic branch, and above with the cell of the 
trichophoric apparatus, by a strand of protoplasm at each end. 
The trichophoric apparatus consists of one or two cells 
according as the total number of cells in the procarp is three 
or four. There is a distinct nucleus in each cell, and the gen- 
eral appearance of the cell contents is very similar to that of 
the carpogenous cell, 2. ¢., the protoplasm containing more oF 
less of the red pigment lies next the cell-wall and encloses 4 
vacuole. In figs. rz, 12 and 13 the cell of the trichophoric 
apparatus is lettered ¢2. The position of the nuclei in the cells 
of the procarps has been shown in many of the figures. In 
some of the specimens (figs. 5—zo) the stain was eosin, in others 
(figs. rr-16) the stain was hematoxylin. 
The structure and development of the trichogyne now remain 
to be considered. This organ is very small and delicate, in this 
species of Ptilota measuring from 40-70, long and 4m wide 1m 
the thinner upper portions. The base of the trichogyne (“‘ car- 
pogonium,” as Schmitz applied the term) is about as wide - 
the cell of the trichophoric apparatus directly under it, but it 
grows narrow very rapidly and runs into the very delicate and 
attenuated upper portion. The base of the trichogyne is not at 
all swollen, nor is there any constriction between it and the uppef 
portion. The cell contents are hyaline in living specimens, and 
quite homogeneous. Stains do not bring out any differentiation 
of the protoplasm aside from a granular structure in the lower 
portion, and the writer has never seen anything that could be 
interpreted as a definite nucleus. 
Such peculiar cytological structure of the trichogy 
merits a farther examination, and the gradual developmen 
withering of the organ will now be described. Starting 
ne cell 
t and 
with 
