364 BOTANICAL GAZETTE [ NOVEMBER 
one of a whorl of small bracts around the cystocarp. These fila- 
ments with the remains of the trichogynes at one side are fre- 
quently met with, and appear in some of the figures illustrating 
the development of the cystocarp. 
We may say at this point that we have never seen any bodies 
attached to the trichogynes that could be identified as anthero- 
zoids. Such observations must be made before the trichogynes 
begin to wither, as then the bacteria and slime put a stop to all 
examination of this point. Sometimes the group of procarps 
contains much foreign matter around the trichogynes, but much 
of the writer’s material was quite clean, and it seemed impossible 
that the presence of antherozoids should escape notice, yet such 
material was covered with developed fruit. 
DEVELOPMENT OF CYSTOCARP. 
We have already stated that the cell at the base of the pair 
of procarps is the carpogenous cell (figs. ¢-7z, ¢). It is very 
curious that the cystocarp should be developed so uniformly 
from a particular cell, and yet this proved true of every spect 
men that the writer examined. This cell is one of the first of 
the cells composing the group of procarps to be formed, and 
consequently is one of the oldest at the time when the cyst0- 
carp begins its development. It is likewise associated with the 
procarp that as a rule is the first of the group to mature. Si 
development of the cystocarp was studied almost exclusively 
from serial sections cut from paraffin, the specimens being 
stained tm toto with Mayer’s acid hamalum, and on the slide with 
eosin. 
The earliest stage of the cystocarp is frequently met 
It consists of a large cell rich in protoplasm, and containing 4 
prominent nucleus, situated in the midst of the group of a 
carps and united to the carpogenous cell of the outer pal : 
glance at fig. 76 will make plain what is meant. The large 
numbered Io is the terminal segment of the procarpic bane 
On the left side of the figure drawn in detail is one of the pro 
carps of the outside pair, and from its carpogenous cell (¢) has 
met with 
