278 REPORT— 1904. 
‘The hind-foot is -relatively more elongated than the fore-foot, and 
measures approximately 0-07 M. in length and 0-038 M. in maximum 
breadth. The first four toes successively increase in size outwards, but 
the fifth is very small—perhaps, indeed, the smallest. The number of 
phalanges is two, three, or four in digits I to III respectively, and 
specimen No. 2 (footprint) shows clearly there are five in digit TV. The 
palm of the foot, so to speak, is of considerable length, and in the hind- 
foot much narrowed posteriorly.’ A small form about 25 mm. in length, 
very closely resembling this, has been noticed from Runcorn on slabs in 
University Museum, Liverpool, and a less perfect one in the Warrington 
Museum from the same place. 
This form differs from D 1 in the greater length of the I digit, in the 
presence of the impression of the palm which is never seen in the im- 
pressions D 1 and 2, in the opposable V digit, and also in the well-marked 
joints of the phalanges. Digits IJ-IV would, however, certainly 
coincide with many of the imperfect D1 prints. This print is fairly 
common in the Keuper in South Staffordshire, though seldom found 
nearly so perfect as those described above. 
Dr. Woodward suggests that these prints may be referred to Rhyncho- 
saurus. He also refers to the slab from the Upper Keuper of Shrewley, 
Warwickshire, described and figured by Murchison and Strickland,' and 
now in the Warwick Museum, as footprints of Rhynchosaurus. This 
slab, owing to its long exposure in the museum, has become deficient in 
the detail, but from a lithograph issued many years ago and from a sketch 
made in 1897 it is clear that it differs from D3. There are nine distinct 
pairs of feet, and in every case the fore and hind feet are side by side, the 
latter, the larger print, being on the outside, whereas in the South Stafford- 
shire example the manus is in a line with and considerably in advance 
of the pes. In the Shrewley slab the digits I-IV of the hind foot are 
distinctly webbed, and there is no trace on any of the impressions of a 
fifth digit. 
D 4. In including these under the Rhynchosauroid prints they may be 
distinguished as D4. Pes only four digits shown, decreasing in size from 
II to [V—all terminated by claws—distinct 
D4, 7—Left pes. mark of webbing connecting the digits claws 
projecting beyond it; total length of print, 
4 cm. ; breadth, 25 mm. 
Manus, three stout toes only shown, with 
indistinct webbing ; total length, 2 em. 
These prints are interesting, as, besides 
giving a complete series, there is a line between 
the rows of impressions the whole length of the 
slab, apparently caused by the dragging of a 
tail. In the forms D1 and 3 the writer has 
failed to see any certain trace of webbing, 
though others have thought they have seen 
indications. It must be borne in mind that 
the example we have from Shrewley is from a bed of sandstone in the 
Upper Keuper, underneath which the Rev. P. B. Brodie has noticed the 
presence of shells of Lamellibranch Mollusca, and in the same bed of 
sandstone casts of Hstheria minutia are frequent ; so that the conditions 
1 Trans. Geol, Society, 2nd series, vol. v. p. 339, plate xxviii. 
