INVESTIGATION OF FAUNA AND FLORA OF TRIAS OF BRITISH ISLES. 279 
were different from those under which the Lower Keuper sandstones were 
formed. 
This form might be compared with Sawrichnites rittlerianus (Hoch- 
stetter), represented by a cast in the British Museum of Natural History 
(R 1474) from the Lower Permian of Semil, Bohemia. The quarry at 
Shrewley has yielded many other prints which still require working out. 
D 5. A print of rather different proportions may conveniently be in- 
cluded in this group, as there is a possibility of its repre- 
senting the corresponding manus to one of the foregoing 
forms. It has not yet been seen in such a position as 
altogether to warrant such an assumption, though it is 
generally associated with prints of D 1. 
In D 5: Only four digits are usually shown, 
No. IV, about 2 cm. in length, being the longest. 
No. I is the shortest, 1 cm. The breadth of IV in the 
widest part is 4 mm. 
The digits are divergent, tapering, and somewhat laterally curved. 
They are in contact at the roots (the articulation with the metacarpals 4), 
which are marked by the presence of round pads. The width of the 
impression here is 15 mm. 
This form will be seen to differ from D 1 not only in its smaller size 
but in its greater proportional breadth and the more rapid taper of the 
digits. 
"E. There is a small footprint described! as E=Rhynchosaurus mint- 
mus (Morton). 
This was first described from fairly perfect examples of both pes and 
manus from Storeton, and afterwards a complete series of 
seven pairs of the same prints was found at Runcorn. E. 2.—Left 
Pes: The outer digit, probably IV, is the longest, * pes. 
10 mm., on the inner side of II, and slightly in the rear is 
a very short hallux. h 
II-IV are divergent, curve inwards, and terminate in 
sharp claws. They frequently have very blunt ends; from 
the centre of each a fine sharp claw projects. There is no 
distinct mark of a claw on the hallux. The breadth of N 
the footprint across the proximal ends of the digits is 
1 em. (pl. iv., fig. 2). 
The manus, represented by four detached toes, is much smaller than 
the pes, being 5 mm. long by 7 mm. broad. The digits are less diver- 
gent than those of the pes ; their diminutive size prevents the recognition 
of any details in a coarse material like sandstone. 
In neither the pes nor manus is there any trace of webbing. On the 
Runcorn slab there are four pairs of impressions of right feet and three 
of left. The impressions of a fore and hind foot are in every case side by 
side, the hind foot being on the outside, with an interval of about a centi- 
metre between it and the manus. The length from one of the feet to 
the next impression in front is 10 cm., and a line drawn outside the hind 
— one side would be 6 cm. distant from a corresponding line outside 
e left. 
This is a very common form, and its size varies greatly, some prints of 
the pes reaching 2 em. in length. The form of the print also varies very 
D5. 1—Manus? 
1 
' Proce. Liv. Geol. Society, vol. vii. 1895, p. 402. 
