— «ties 
TRANSACTIONS OF SECTION D. 593 
One point, however, remains to be cleared up, and that is, the absence of 
browas in F, and F,, these being in all cases apparently displaced by greys. 
4. Dutch Markings.—In F, the offspring from one albino were, as a 
rule, uniform in colour, like the coloured parent, while those from the other 
albino were all more or less marked with white on the fore extremities. In F, 
and F, the uniform hybrids were, as a rule, constant, while the marked ones 
produced a proportion of true Dutch-marked rabbits, as well as the ordinary- 
marked and some uniform ones. 
These results suggest that one albino contained the factor for Dutch markings 
while the other albino did not. Experiments are now in progress to work out 
this interesting question. 
4. Huperiments on Heredity in Fowls. By R. C. Punnerr, 
5. An ‘Intermediate’ Hybrid in Wheat. By R. H. Brirren, 
In the majority of cases investigated up to the present one character of agiven 
pair (the dominant) masks the other (recessive) in the first generation (F,). 
In the case of Triticum Polonicum (Polish wheat) x T. turgidum (Rivet 
wheat) and its reciprocal, the hybrid does not show this sharp separation of 
dominant and recessive characters, it being intermediate between the parents 
with regard to certain pairs. Thus in Polish wheat the glumes, grain, and inter- 
nodes of the spike are long, and it ripens early; in Rivet wheat the glumes, grain, 
and internodes are short and it ripens late; whilst the hybrid has an intermediate 
length of glumes, grains, and internodes, and also ripening period. The hybrid 
has therefore a distinct character of its own in which one cannot recognise definite 
dominant ard recessive characters. However, in the following generation (F,) 
the splitting is of the type which Mendel’s work has made so familiar to us. We 
find long, intermediate, and short glumes, early, intermediate, and late ripening, 
&c., in the proportion of 1 : 2 : 1, showing that, in spite of the fact that there is 
no marked distinction into the usual dominant and recessive characters, the 
gametes are still pure with respect to the characters they carry. 
6. Laperiments on the Behaviour of Differentiating Colowr-characters 
in Maize. By R. H. Lock, B.A. 
Maize as grown in Ceylon is of a flint variety, but shows a complex mixture 
of the colours white, yellow, blue, and red. 
White, yellow, and blue grains occur mixed in the same cob. The red colour 
appears either in all the grains of a cob or not at all. 
The problem attempted was to discover, by the method ot growing the off- 
spring obtained by definite pollination, how far these several colour-characters 
conform to Mendel’s law. 
When a few yellow grains appear in an otherwise white cob such grains must 
be supposed to be the result of accidental impregnation of pollen bearing the 
yellow character. On sowing these grains and fertilising the female flowers with 
the pollen of a white variety the plants yield 50 per cent. of yellow and 50 per 
cent. of white grains within the limits of error due to the size of the samples 
afforded by the cobs, which contain as a rule from 300 to 800 grains. 
Blue grains, appearing in the same way in an otherwise white cob, when 
treated as above yielded, as a rule, a much smaller percentage of blue grains—e.g., ° 
30 per cent.—and the proportion is also more variable. The intensity of the blue 
colour also varies considerably. But on sowing the grains from such a cob, 2.e., 
one with 80 per cent. of blue grains, and pollinating with white, it was found 
that the blue colour reappeared in the offspring of approximately 50 per cent. of 
the original grains; i.e., in those of the 30 blue grains, and in those of 20 out of 
1904, QQ 
