386 BOTANICAL GAZETTE [ NOVEMBER 
said to take place in the pollen mother cell. So it has been described for 
Lilium Martagon by Strasburger, Guignard, Farmer, and Miss Sargant; also 
for Larix Europea and other plants. In all these cases, too, the chromosomes 
are distributed by means of two longitudinal fissions; while in A//ium fistu- 
fosum their distribution in the second division of the pollen mother cell is 
effected by transverse fission. It will be noted that in so far as the reduction 
in Allium fistulosum deviates from the ordinary plant type it approaches what 
has been reported in certain animals, especially in some copepods and 
Gryllotalpa, where the second division of the spermatocytes is attended by 
transverse splitting of the chromosomes, and the reduced number of chromo- 
somes appears early, even asearly in Cyclops brevicornis, according to Haecker, 
as the primordial cells of the blastula stage. 
Transverse fission of chromosomes in plants has been reported by Calkins 
in Pteris and Adiantum, by Mottier in Lilium, and by Schaffner in Lilium 
Philadelphicum.—W.R.S. 
