1898 } THE DEVELOPMENT OF SOME ANTHRACNOSES 113 
which are common to Gleeosporium, Colletotrichum, Volutella, 
and Vermicularia is not a constant character, but may be absent 
throughout the entire cycle of development of a species, or they 
may be forced in these same species by a lack of nourishment. 
The presence of a perithecial stage has been proven in four dif- 
ferent species, of which the host plants vary widely in habits and 
structure. One of these exists as a saprophyte in a natural 
state (the one in connection with the vanilla anthracnose is 
omitted), while the other three are saprophytes in artificial cul- 
tures; whether they occur in nature has not been determined. 
Two of the perithecial forms were connected with species of 
the genus Colletotrichum, and two with Gleeosporium.” 
The colonies produced in the species with which perfect 
forms have been connected agree in producing a loose open 
growth about the center, but all show specific differences in for- 
mation of stroma, pigment, or arrangement of fruiting sori. 
While the conidial forms show greater variations in structure 
than do the perithecial stages of the different species, the growth 
characters of the colonies from ascospores resemble those of the 
conidial stage with which they are connected more closely than 
they resemble each other. 
The perfect forms approach the genus Gnomoniella, agree- 
ing in the submembranaceous, subglokose, subcutaneous-erum- 
pent perithecia; in the cylindrical, clavate, 8-spored asci; and 
the continuous hyaline conidia. They differ in the curved 
conidia, which in Gnomoniella are typically ovate, oblong sub- 
filiform and straight, although in species of Gnomoniella, G. 
amena (Nees) Sacc. and G. fasciculata (Fckl.) Sace. they are 
curved. The genus under consideration does not show ne 
slender, somewhat elongated beaks found in Gnomoniella, which 
are surrounded at the base with a white, cleft collar formed of 
the ruptured epidermis of the host, and the necks are hairy, 
while in Gnomoniella they are smooth. In shape the perithecia 
resemble the genus Camptospheria, but they differ from this 
* The presence of sete in the cultures has been so variable as to — 
tion whether they form a well-founded basis for distinguishing these two genera. 
