302 BOTANICAL GAZETTE [NOVEMBER 
pistils are not entirely closed until a very short time before 
fertilization, and then in the same manner. The further devel- 
opment of the pistil after fertilization is, however, a little dif- 
ferent in the two genera. In Myosurus, instead of enlarging 
rapidly so as to leave a large cavity which the ovule fills only 
in part, the pistil enlarges only as does the ovule, leaving no 
cavity. The walls, too, do not become stony, thus allowing 
(what is very difficult in Ranunculus) the study of the devel- 
opment of the embryo. 
ANEMONE. In the first stages of the development of the 
pistils this genus resembles Myosurus very closely. In fact, the 
development throughout of the single large ovule is as in that 
genus (figs. 20-23, 26). A slight difference in the shape of 
the pistil is noticeable, in that the cavity is prolonged somewhat 
above the ovule. In Anemone Caroliniana, after the ovule has 
curved down into the cavity of the pistil, there appear on the 
edges of the laminae, which are now closing together, two pro- 
jections. These increase in size, growing down into the cavity 
of the pistil above the first ovule. They remain merely few- 
celled papillae in this species, one on each lamina, the central 
cells resembling archesporial cells (fig. 2¢). This occurs also 
in other species of this genus, the papillae in some species often 
becoming well-marked, rudimentary ovules. This isso common 
that Baillon describes? Anemone as being provided with five 
ovules, four being aborted and one descending and fertile. 
CLEMATIS LIGUSTICIFOLIA. The pistils in this species are even 
more elongated than in Anemone, and have the cavity corre- 
spondingly elongated above the ovule. Unfortunately, it was 
impossible to obtain the younger stages, but the close agree- 
ment of the older stages with those found in Anemone makes 
it practically certain that the course of development is the 
same. The mature ovule is situated exactly as in Myosurus and 
Anemone. Like the latter, however, on each side above the 
large ovule the lamina bears one or two rudimentary ovules 
which project into the space above it (fig. 25). The only dif- 
*BAILLON: Mémoire sur la famille des Renonculacées. Adansonia 4:50. 1864. 
