306 BOTANICAL GAZETTE | NOVEMBER 
only as the pistil grows that the funiculus also increases in 
length. 
ROSACE., 
This family contains representatives of many types of flower- 
structure. Of these types the Potentillee have been regarded 
hitherto as the simplest. In this tribe the pistils are very numer- 
ous, on a rounded receptacle, which is expanded below into a 
shallow cup, on whose edge are borne the numerous free stamens, 
the petals and the sepals. In Fragaria and Potentilla each pistil 
is uniovulate, while in Geum it is biovulate. As a rule through- 
out the family the pistils are biovulate, and in some genera even 
multiovulate. The only genera studied as representatives of 
this family were Potentilla and Fragaria, it being the aim to 
determine whether the remarkable similarity that these show to 
Ranunculus is also found in the processes of development of the 
parts of the flower. 
POTENTILLA Monspe.iensis L. The pistils first appear, as in 
Ranunculus, as small papillae on the surface of the pistil-bearing 
part of the receptacle. The first to appear are at the base and 
the others arise successively towards the top of the receptacle 
(jig. 45). As the pistils enlarge they become hollowed out 
above. A comparatively small opening is produced on the 
upper side of the pistil (fg. 46), which is made still narrower by 
the thickening of the edges of the laminae for about half the 
distance from base to apex. This thickening is sometimes 
accompanied by a more active growth in width of that part of 
each lamina, so that viewed from the side it appears as a rounded 
lobe, as shown in fig. 47, where the dotted line shows the 
more usual form. From one of these thickened edges or lobes 
a small papilla begins to grow inward and downward, later turn- 
ing upward again. This is the ovule. It is at first lateral in its 
position, but the lamina to which it is attached grows more 
rapidly than the part opposite, so that the ovule finally occupies 
a median position (fig. 48). The ovule, when the pistil is ready 
for pollination, is anatropous, with the funiculus on the ventral 
