350 BOTANICAL GAZETTE [ NOVEMBER 
turning over of the edges (figs. g-8). The cotyledons are separated 
for some distance near the tops of the lobes. In Juglans the cotyle- 
dons are deeply two-parted (fg. 7); the cotyledon in the nut being 
U-shaped (fig. 3) and their lobes united to the summit (fg. 2). In 
Hicoria the shape of the embryo varies greatly in the different species. 
Hicorta_glabra and Hicoria microcarpa, which in this region are not 
sharply separated, have the embryos much alike, but in Aicoria glabra, 
the division between the cotyledons is twice as deep as in Aicoria 
microcarpa. 
_ In germinating, the seed splits from the micropyle and the tip of 
the radicle and the two or four basal lobes of the cotyledons push out 
together. The petioles of the cotyledons lengthen and carry the 
plumule out of the nut. The cotyledons remain in the nut and do 
not decrease to any appreciable extent in size, but become very rancid 
in taste and are filled with a yellowish oil similar to that found in the 
husks of walnuts and butternuts. The valves of the nut usually remain 
slightly connected at the hilum but often are split entirely apart, and 
in one specimen of Juglans cinerea where the nut was on the surface 
of the ground, the two valves had separated and lay one on each 
side of the stem. The cotyledons, where exposed to the light, were 
green and could be easily drawn from the nut. 
The root of the seedling becomes greatly thickened. In one 
specimen of Hicoria glabra, it was 6™ long and 1.3™ in diameter, and 
others were nearly as large. The outer portion becomes brown and 
fissured, the fissures extending to the endodermis. In older seedlings 
ment. As the seedlings naturally germinate under trees where ” 
nuts would be buried under leaves, and as the internodes have not - 
power to lengthen much, Lubbock’ suggests that the first leaves - 
5 LuBBocK, SiR J., of. cit. 
