1909] OTTLEY—JUNIPERUS 35 
the generative cell. This corresponds with the condition at this time 
in Thuja as described by LAND (14). As noted by Nor&n, no starch 
appears in the pollen tube at this time. Previous to the division of 
the generative cell its cytoplasm becomes decidedly alveolar in appear- 
ance (jig. 13), suggesting the appearance of this cell in Zamia (WEBER 
29), Cycas (IKENO 9), and Sequoia (LAWSON 15). Nor&n does not 
speak of this appearance, but describes the cytoplasm as coarse 
grained and containing a large amount of finely divided starch which 
might easily escape observation. I was unable to find any signs of 
starch in this cell at any time in its history. 
Just before fertilization the generative cell divides and forms two 
cells equal in size (jig. 27), as shown and described by Nor&n. This 
agrees with Biota (STRASBURGER 27), Thuja (LAND 14), Taxodium 
(COKER 5), and Cryptomeria and Sequoia (LAWSON 15, 16). When 
mature the two sperm cells in Juniperus are hemispherical in shape 
and lie with their flat sides face to face, but not in contact, as described 
by Norén, and in Thuja by Lanp (14). The division of the genera- 
tive cell occurs normally in Juniperus after the tube has reached the 
neck of the archegonium, but in one instance two similar nearly 
spherical cells which resembled generative cells appeared in the pollen 
tube when it was about half-way through the nucellus. 
It would seem that the two sperm cells may both be capable of 
functioning, as LAND (14) thinks is the case in Thuja. They are 
equal in size, the archegonia are borne in complexes, and the tip of 
the pollen tube is pressed against the neck cells of several archegonia, 
thus making it possible for the sperm cells from one tube to enter 
different archegonia. Two or three pollen tubes may reach the same 
archegonial complex, and there is very little branching of the tubes 
in their way through the nucellus. 
FEMALE G TE 
On March 28 the ovuliferous cones had appeared, but there were 
no indications of ovules. The first stages in their development were 
observed May 1. There are, as a rule, three ovules borne in the same 
horizontal plane at the apex of the cone. The integument arises as a 
little Swelling at the base of the nucellus, and by May 8 it had passed 
beyond the top of the nucellus. But one integument is present, and 
