1909] ; OTTLEY—JUNIPERUS 39. 
but cannot explain them. Fig. 26 shows these radiations in an early 
prophase of the division of the central cell. 
After the ventral canal nucleus is cut off, it moves to the micropylar 
end of the cytoplasm and usually disorganizes before the pollen tube 
bursts. Fig. 28 shows an unusually large ventral canal nucleus. 
The egg nucleus increases in size and moves slowly toward the central 
vacuole, taking up a position just above it. It usually possesses a 
beautiful chromatin network and a large nucleolus or several smaller 
nucleoli. At this time both the radiations and the protein vacuoles 
disappear, and many darkly staining specks appear in the egg cyto- 
plasm (jig. 27). Contrary to what I have observed and StupsKy 
(22) has reported, Nor&n (20) says that these centers of radiations, 
so noticeable at the time of the division of the central cell, increase 
in size during the maturation of the egg. There is no evidence what-. 
soever of this in the material which formed the basis of the present 
study. 
The archegonia form a complex which is surrounded by a layer 
of sheath cells. These are small and filled with deeply staining 
protoplasm. HormeIsTER (8) says that Juniperus frequently has 
archegonia in abnormal positions and Nor&én has made the same 
observation. The only case of archegonia in abnormal positions 
that was observed, is shown in fig. 36. Here the archegonium is out- 
side of the layer of sheath cells, but in other respects it appears normal. 
FERTILIZATION 
Fertilization follows directly upon the division of the generative 
cell and the maturation of the egg cell. In the summer of 1905 
fertilization in J. communis occurred on June 17, 20, and 21. In 
ovules put up on the same day were found divisions of the central 
cell to form the egg cell and ventral canal nucleus, undivided genera- 
tive cells, sperm cells, fertilization, and proembryos. This would 
indicate that these divisions take place very rapidly. A mass of 
densely staining, coarse-grained cytoplasm accompanies the sperm 
nucleus on its entrance into the egg. Whether the other contents 
of the tube enter the archegonium was not clearly determined; in 
one preparation there were densely staining bodies in the upper part 
of the archegonium, which might be the disorganized remains of the 
