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1909] PACE—GAMETOPHYTES OF CALOPOGON 127 
The other ovules were in the mother cell stage (fig. 4). The integu- 
ment is only a trifle niore advanced in the latter than in the former, 
but the nuclei are in all stages of synapsis. The archesporial cell 
may become the mother cell without division, and no parietal cell was 
ever found. But it is possible that in some cases it divides; for an 
archesporium of more than one cell was rare, being found only once 
(fig. 26), and yet several ovules with two mother cells in position to 
have been formed by division of a single archesporial cell were found 
(figs. 27, 29). It is evident that these must have originated by a 
division of the archesporial cell, or the archesporium must have 
consisted of two cells. 
A more remarkable case still is that in which two distinct sporo- 
genous areas are differentiated in the same ovule (figs. 28-31). Fig. 31 
shows several cells of nucellar tissue between the two mother cells, 
the cut being across the ovule in a different direction from that of 
jig. 28. A somewhat later stage, similar to fig. 28, is shown in fig. 30. 
Fig. 32 shows at least one of the mother cells divided, and one of the 
resulting daughter cells increasing for the second division, the other 
disintegrating in the usual fashion. The presence of more than one 
mother cell is not rare in my material. Of about sixty ovaries cut, 
thirteen showed ovules with two mother cells, or stages derived from 
this condition. In these thirteen ovaries, the ovules showing this con- 
dition vary from one to seven, giving a total of thirty-seven. Of these 
thirty-seven, twenty-one are similar to figs. 28 and 37, and sixteen 
resemble jigs. 27 and 29. There was in all probability double this 
number, for only those were counted in which both cells appeared 
in the same section. No attempt was made to trace the ovules from 
section to section for this condition, and it is evident that there are 
more chances against getting both in the same section than in favor 
ofit. Fig. 3risa drawing from a single section, but the whole ovule 
was traced carefully from section to section for evidence of the coales- 
cence of two ovules. The only abnormal appearance was the unusu- 
ally broad funiculus. Fig. 33 shows the megaspores derived from 
the two adjacent mother cells. In each case the chalazal megaspore 
is developing and the other three are disintegrating. In the lower 
or micropylar group the megaspores are in a row—the usual row of 
four, except that the wall is lacking between the two upper megaspore 
