= CRs o 2 en ee ay SY. 
3 Ig09] BROW N—EMBRYO SAC OF HABENARIA 243 
antipodal, while the other spindle forms twoantipodals. Walls cutting 
off the egg, synergids, and antipodals are formed on fibers connecting 
the nuclei. Owing to their position nearer the center of the sac, 
the polar nuclei are left free in the cytoplasm (fig. 5), as has been 
pointed out by STRASBURGER (?05) for Drimys Winteri. After their 
formation, the egg and synergids enlarge considerably, while the 
antipodals soon degenerate ( jig. 6). When the egg and synergids 
have about reached their mature size, the polar nuclei fuse (fig. 6). 
No cases were observed in which the polar nuclei were in con- 
Fics. 8-1o. Older embryos: s, suspensor; e, embryo. 
tact without fusing, as have been described for other forms by 
STRASBURGER (700) and Nawascutn (’00). The pollen tube comes 
through the micropyle and discharges the two male nuclei into the 
embryo sac. One of these fuses with the egg, while the other fuses 
with the product of the fusion of the polar nuclei to form the primary 
endosperm nucleus. This enlarges somewhat, but without dividing 
begins to degenerate at about the time of the first division of the fer- 
tilized ege (fig. 7). ee 
The first division of the fertilized egg is transverse to the longitudi- 
Nal axis of the embryo sac. Of the two resulting cells the chalazal one 
