44 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 8o 



Matthew ' as Dromillopiis quadrifidus from the Coal Measures of 

 Joggins, Nova Scotia. The presence of a distinct tail drag, and its 

 absence in the Joggins trackway (compare figs. 19 and 20), differ- 

 ences found in the structure of the forefeet, longer stride, and 

 greater width of trackway all point to its specific distinctness from 

 the Nova Scotian species ; hence the specific name parvus is proposed 

 for its reception. Hind foot: Length 9 mm., width 8 mm. Four digits 

 of which the outer is set ofif from the other three. These are long 

 and slender, regularly increasing in length toward the outside of 

 the foot, the third being the longest, the fourth considerably shorter. 



# 



"^ 'M 





i" 



Fig. 20. — Dromillopiis quadrifidus Matthew. Type. Diagram of 

 trackway. About natural size. (After Matthew.) 



The sole is well impressed and the full rounded outline of the heel 

 is shown in figure 19, whereas in the Joggins species the sole is 

 scarcely distinguishable. The foot is shown to have been semiplanti- 

 grade, not digitigrade as originally characterized by ^Matthew. Fore- 

 foot: Length about 8 mm., width 8 mm. While none of the so-called 

 hindfoot impressions show the detailed structure plainly, it is clearly 

 evident there were only four toes on the manus. The sole seems to be 

 more broadly rounded than in the pes. A description of the other 

 details of the foot must await the discovery of better preserved 

 specimens. 



The imprints connected by dotted line in figure 20 were regarded 

 by Matthew as having been made by the hindfeet. No reasons were 



' Proc. Trans. Roy. Soc. Canada, Vol. 10, 1904, p. 91. 



