48 The Botanical Gazette. (February, 
The zoospores are peculiar in appearance and can readily 
be distinguished from any similar bodies known to the writer 
by the characteristic disposition of their contents. In general 
form they are oval or elliptical and are, at least in many 
cases, biciliate; the two cilia arising side by side from the 
smaller end of the spore. In some instances it has been 
found impossible to make out more than a single cilium even 
after the zoospore was stained; but the presence of two (fig. 
11) has been determined definitely in so many cases that this 
number may be considered as typical. The nucleus is very 
large and sub-triangular in outline, its base connected with that 
of the cilia by a fine strand of granular protoplasm. In front 
of the nucleus lies a broad and distinct mass of granular pro- 
toplasm while small groups of granules occur here and there 
around it in the otherwise nearly clear cytoplasm. In gen- 
eral appearance they are not unlike the zoospore of Gona- 
podya, but may always be distinguished by the form and po- 
sition of the nucleus and the evident connection of the lat- 
ter with the base of the cilia. As the fungus developes, 
growing as it almost invariably does in tufts, it forms the cen- 
ter of a dense mass of bacteria which finally choke the spo- 
rangia completely; so that as arule only those first formed are 
able to discharge their contents. Asa result the zoospores 
commonly die without escaping; the remains of their large 
nuclei just mentioned filling the sporangia, as is indicated 
in the central sporangium of fig. 3. These dead nuclei were 
taken by Reinsch for the zoospores themselves and are repre- 
sented in his plate xvI, fig. 8. 
Associated with the sporangia are often found numerous 
slender filaments which arise as buds ina fashion exactly sim- 
ilar to that by which the former are produced. They are 
very slender, simple or irregularly branched, without septa, 
and often greatly exceed the sporangiain length (fig. 2). In 
a majority of cases, however, they are wholly absent oF 
undeveloped (fig. 1 at the right) and they seem to bear no 
definite relation either to the sporangia or to the resting 
conidia. They seem to be quite sterile and although Reinsch 
suggests that they may be antheridia they are probably with- 
out definite function, and are certainly not male organs. 
The resting spores, already referred to, and described by 
Reinsch as doubtful oospores, do not as a rule make their ap- 
pearance until some time after the sporangia have been de- 
