1896. ] Development of Champia. I15 
The development of the favellz of spores now remains to be 
considered. We must start with stages such as are shown in 
gs.9 and 10. It will be remembered that at this stage the 
trichogyne and carpogonium have entirely disappeared so that 
the cell lettered » in these two figures is the one directly under 
the carpogonium. This cell gives rise to the cystocarp. By 
several transverse divisions, which are usually somewhat ob- 
ligue, it forms a short branch consisting of four or five cells, two 
stages of which will be seen in figs. 11 and 12. The branch 
is greatly bent to one side in later stages and then oblique 
walls are formed across the upper segments in a very irregu- 
lar manner, and there results a compact mass of cells divided 
"Pinto a number of lobes. The entire young cystocarp is 
really a very complex set of filaments consisting of angular 
and irregularly shaped cells, but this structure is not readily 
‘town in sections. The cells at first contain each a single 
tucleus, but when the Cystocarp is mature it is apparent that 
the spores are borne on the ends of short branches (fig. 14) 
a cells are multinucleate although each spore is uninu- 
ate. 
. As the Cystocarp arises from the cell of the procarpic branch 
Just below the carpogonium, this is the place where we should 
“pect to find cross fusion with auxiliary cells if such exists. 
The writer has observed no specimens which would indicate 
- an auxiliary cell ever completely fused with any of the 
; . rectly concerned with the development of the cysto- 
re ‘ae, thing was ever seen that could be sca eo ae 
cells “entral cell” of Hauptfleisch. It is true that auxiliary 
ana af¢ Clustered around the cells of the young cystocarp, 
, it would be very easy for them to unite by protoplasmic 
processes, but the writer saw no clear evidence of such cyto- 
¢ fusion, much less of any nuclear disturbances. In 
Me cases the cells of young cystocarps have been quite 
ed. _Dlasmic fusion. es ie 
at Is the ¢ l cytoplasmic union 0 
th ause of the very general cytop c 
ens at the base of the cystocarp? The phenomenon be 
S after the trichogyne has disappeared and the gerd 
Presumably been fertilized. It is concerned with the 
