124 The Botanical Gazette. [Marc 
F. C. Newcombe, to whom I here express my sincere thanks 
for kind assistance rendered in various ways. 
Development of the embryo-sac. 
The young ovaries arise as protuberances around the edge 
of the flattened receptacle, and soon there appears in the it- 
terior of each one a schizogenetic cleft on the inner side o 
which the nucellus is formed. This cleft increases in size quite 
rapidly by enlargement and division of the cells of its walls 
thus making room for the developing nucellus. | When the 
nucellus has obtained some considerable size, there can be 
seen in it a large hypodermal cell which appears to be the 
archesporium. No division of this cell into two was observed 
but at a later stage the large macrospore shows the remaili 
of a former cell at its micropylar end (fig 1.) which is the 
tapetal cell. The ovule soon becomes anatropous. The I 
teguments are two in number, but on the side of the funice 
lus the outer one is generally not developed. As the embry 
Sac increases in size the ordinary divisions of its nucleus take 
place. First it divides into two, one of the daughter 1 
passing to the upper and the other to the lower end of the 
sac, after which each of these, by two successive divisions — 
produces four nuclei, thus making the typical eight-celle 
be 
large nucleolus, but sometimes two (fig. 2). The two cel 
gid lie side by side. They are surrounded by granular 
ak 
They stretch across the entire upper end of the sac, and 
beneath them the oosphere is suspended in a dense Be- 
cytoplasm. Its nucleus is usually ellipsoidal in shape 
yond the oosphere and lying free in the cytoplasm is the ef 
per polar nucleus. Its centrospheres usually lie om we 
arrangement in the €gg-apparatus. There are two sa 
