38 BOTANICAL GAZETTE [yuL 
are present, but many staminodia are inserted near the base of the 
ovaries. 
Ovary.—In the youngest material examined, the beginning of 
the flower appears as a Slight protuberance on the surface of the 
spadix. A transverse crease soon appears, dividing this into two 
ill-defined lobes (fig. 13) which are destined to develop into the two 
carpels, and between the carpels appears the placenta, representing 
the axis of the flower (fig. 14). The carpels rapidly increase in size 
and overtop the placenta (figs. 15, 16). At the same time the ovules 
appear as lateral outgrowths of the placenta, filling the two cavities 
formed by the growth of the carpels (fig. 75).. The continued’ growth 
of the carpels causes them to approach each other until there is left 
between them only a narrow canal, which divides over the apex of 
the placenta in such a way that the two branches lead down into 
the cavities of the carpels (fig. 77). On the upper and inner sur- 
face of each carpel, and leading down to the stylar canal, occurs a 
circular patch of somewhat viscid stigmatic cells (fig. 17). 
Ovutr.—The ovule first appears as a dome-shaped mass of 
undifferentiated cells projecting laterally into the carpellary cavity 
(fig. 15). The inner integument soon appears as an ill-defined ring 
about the upper portion of the ovule, and by the time the carpel has 
partially covered the placenta, the integument has grown out even 
with the apex of the nucellus, and the outer integument has begun 
to appear (fig. 16). At this stage the ovule is orthotropous. The 
lower surface now ceases‘to develop, while the upper continues to 
grow, thus finally making the ovule anatropous. At the same time, 
the integuments lengthen greatly, the outer one much exceeding the 
inner one (jig. 17). 
ARCHESPORIUM.—Before the ovule becomes anatropous, the 
archesporial cell may be recognized by its greater size and by its 
nucleus (fig. 19). Before the first division of the archesporium the 
overlying epidermal cells usually divide once (fig. 20). The arche- 
sporial cell, which in this case is the mother cell, gives rise to four 
megaspores (jig. 21), the outermost one of which produces the em- 
ryo sac, 
_ Empryo sac.—The functioning megaspore increases greatly in 
size, destroying as usual the other megaspores and some of the 
