THE EMBRYO SAC AND EMBRYO OF GNETUM GNEMON 
CONTRIBUTIONS FROM THE HULL BOTANICAL LABORATORY I12 
Joun M. COULTER 
(WITH PLATE VII) 
Gnetum Gnemon has been made conspicuous among the other 
species of Gnetum chiefly by the investigation of Lorsy published 
in 1899. The structures described were of such interest that it 
seemed desirable to supplement the somewhat incomplete account 
by a further examination. Accordingly material was obtained from 
the Philippine Islands and from British Guiana; the former from 
Dr. H. N. Wuuirrorp, and the latter from Mr. A. W. BaRtLert, 
director of the Botanic Garden at Georgetown. This material was 
first assembled by Dr. W. J. G. Lanp, of this laboratory, in connec- 
tion with his investigation of Ephedra; and he has kindly turned it 
over to me for separate study. The preparations and drawings were 
made by Dr. SHicfo Yamanoucui, of this laboratory, and to his 
technical skill the results are largely due. 
The material included stages from two successive seasons, but 
unfortunately many intervening stages were not represented, so 
that no continuous account can be given. However, certain facts 
have been discovered that supplement and correct the previous 
accounts. 
EMBRYO SAC 
_ Lorsy described the embryo sac of G. Gnemon as showing an 
interesting deviation from those found by KarsTEN? in other species 
of Gnetum. Instead of containing only free nuclei at the fertiliza- 
tion stage, the embryo sac of G. Gnemon was described as containing 
Tsy, J. P., Contributions to the life history of the ean Gnetum. Ann. 
Fatal he Wittens II. 1:46-114. pls. 2-11. 1899. 
2 KarsTEN, H., Bot. Zeit. 50:205-215, 221-231, 237-246. pls. 4,6. Roxas Ann, 
Jard. Bot. Buitenzorg 11:195-218. pls. 17-19. 1893; Cohn’s Beitr. Biol Pflanzen 6: 
337-382. pls. 8-11. 1893. 
43] (Botanical Gazette, vol. 46 
