1908] BURLINGAME—PODOCARPUS I7I 
remains large and the cell itself is often much larger than when 
two are cut off (ext jig. 1). Sometimes the three resulting nuclei 
then lie in a row across the spore (text fig. 1) and differ but little from 
one another in appearance. The walls separating them then run 
nearly straight across, and it is only by courtesy that the tube nucleus 
can be said to be free in the spore cytoplasm. To all intents and 
purposes it is as much walled-off into a cell of its own as the pro- 
thallial nucleus, for both eventually are freed in a common cytoplasm 
(text fig. 2). Further, it is to be noted that in this case the prothallial 
cell does not have a fixed position with respect to the wings, as in the 
ordinary course of events. 
After the tube nucleus has been cut off, the primary sperma- 
togenous cell divides transversely, giving rise to two cells (jig. 25). 
sually these cells differ markedly in size, the smaller lying to one 
side and the larger lying nearly in the center of the pollen grain (fig. 
26). In this case one may safely use the usual terms for them and 
speak of the larger as the body cell and the smaller as the stalk cell. 
It not infrequently happens that the two derivatives of the generative 
cell are about equal in size (figs. 24, 27; text figs. 5, 6), though even 
then one is centrally placed and the other laterally. Before the 
division of the generative cell, the second primary prothallial cell has 
usually divided at least once. In this case the generative cell often 
sinks down among its derivatives (figs. 23,27). The stalk cell is some- 
times placed transversely to the body cell (text figs. 5, 6). Whether 
these large stalk cells will produce male cells or not can only be con- 
jectured, though the fact that some of them seem to retain their 
‘ytoplasmic envelope, almost as distinct from the cytoplasm as the 
body cell itself, up to a certain early stage, would lead one to sa ort 
that in some cases they might do so, Text fig. 7 may show three deriv- 
atives of the generative cell and two prothallial cells in addition to 
the tube nucleus. One of the three derivatives of the generative 
cell has no cytoplasmic sheath and is probably a stalk nucleus; the 
‘Wo lying close together are ensheathed, though I have been unable 
to ascertain definitely whether each has a distinct sheath or whether 
th lie in a common sheath, or whether perhaps one of them is 
not merely beneath the cytoplasmic sheath of the other. If either the 
St or second supposition is true, there still remains the question 
