1908] BURLINGAME—PODOCARPUS 173 
stages shown in the plates (jigs. 26, 27; text fig. 7) occurred along 
with the youngest. In some cases the cell walls were still intact, 
while in other pollen grains not more advanced, the nuclei and body 
cell were free in the common cytoplasm (text figs. 1, 3, 7). 
Discussion 
The facts presented in the preceding account are in the main 
confirmatory of those already reported by CoKER and by JEFFREY 
and CHRYSLER for other species of Podocarpus; hence their main 
value is that they offer additional evidence that the phenomena 
described by these authors are not limited to the species studied by 
them. These authors and others as well have called attention to 
the resemblances to the Abietineae in the winged pollen, thick sporan- 
gium wall, and general similarity of the staminate strobilus. 
The main interest centers’ about the development of the male 
gametophyte. CoKER (2) reported two primary prothallial cells 
in P, coriacea, which divide amitotically. JeFFREY and CHRYSLER 
(5) have recorded as many as eight in the species studied by them. 
Miss Younc (14) has also recorded as many as four in species of 
Dacrydium. THompson (13) has found 30 to 4o prothallial cells 
In Agathis; and LopriorE (8) has reported a cell complex in Arau- 
caria that reaches about 1 5 in number before the walls break down, 
When the freed nuclei may then further divide until as many as 
44 may be found in the pollen tube. He interprets these as sperma- 
togenous cells and cites the case of Cupressus as analogous. CHAM- 
BERLAIN (2) has already called attention to the fact that the drawings 
seem to indicate that these nuclei are really prothallial and that 
there is only one body cell. In this view Miss Younc (14) has con- 
Curred, as it would seem most students of gymnosperms must. RE- 
NAULT (10, rz) and OLIVER (9) have described the multicellular 
Pollen grains of the Cordaitales and Stephanospermum, where as 
many as 20 cells are found in the male gametophyte; whether they 
are prothallial or spermatogenous is uncertain. 
Catpwext (1) has reported as many as nine or ten body cells in 
Microcycas, along with a single stalk and single prothallial cell. 
CHAMBERLATN in a forthcoming paper reports that Ceratozamia 
®ceasionally has four sperms. JUEL (6) found a variable number 
