THE MORPHOLOGY OF PHYLLOCLADUS ALPINUS 
CONTRIBUTIONS FROM THE HULL BOTANICAL LABORATORY 118 
N. JOHANNA KILDAHL 
(WITH PLATES XX—XXII) 
The only account of the morphology of Phyllocladus is that of 
Miss AGNes RoBERTSON (18) in 1906, and this deals exclusively with 
the anatomical structures and with the affinities of the genus. 
The material for the present study was collected by Dr. L. Cocx- 
AYNE of New Zealand. It consisted of two collections: one of ovu- 
late and staminate strobili, collected November 1, 1906; the other 
of ovulate strobili, collected December 11, 1906, from cultivated 
trees. Unfortunately, the two collections did not afford a very close 
series, and it was impossible to get at some of the most important 
points, as the development of the microsporangia, microspores, pollen 
tubes, female gametophyte, archegonial neck cells, ventral cell or 
nucleus, and embryo. Dr. CocKAYyNE is making collections of the 
lacking stages at the present time, and it is hoped that a complete 
description can be presented later. Some material was also obtained 
from Cambridge, Mass., through the kindness of Dr. B. L. Rosrnson, 
Consisting of three ovulate cones, taken from the Gray Herbarium 
and collected on the Wilkes expedition in 1838-1842. At what time 
of the year these cones were collected is not known, but they showed 
Somewhat later stages than those of the last collection made by Dr. 
Cockayne on December 1 1. It is needless to say that this herbarium 
Material was very much shrunken, and it was impossible to obtain 
from it any definite structure. 
Dr. Cockayne’s materia] was killed and fixed in the field, in 
7° per cent. alcohol and formalin; cut 5 “and 7 w thick; and stained 
Part in safranin and gentian violet, and part in iron alum. 
Staminate strobilus 
The staminate strobili are formed laterally, in clusters of 2-8 at 
the tips of the lateral branches. Miss ROBERTSON (18) reports one 
Staminate strobilus bearing an ovule at its base, which may indicate 
339] [Botanical Gazette, vol. 46 
