396 BOTANICAL GAZETTE [NOVEMBER 
fact that a protostelic genus Lygodium occurs among the living Schizaeaceae, 
throws no light on the morphological nature of the pith in the central cylinder 
of the other schizaeaceous genera Schizaea, Aneimia, and Mohria.—E. C. 
JEFFREY. 
Mitosis in Cynomorium.—Baccarini"® has published an account of vegeta- 
tive mitosis in Cynomorium coccineum (Balanophoraceae). From a study of the 
nucleus in meristem of the roots and in parenchyma of the stem, he finds the 
following stages: (1) Prophase: (a) chromatin granules are uniformly distributed 
throughout the fundamental mass of the nucleus, with or without still larger 
chromatin knots or joints, the chromocenters; (6) the chromatic granules, separat- 
ing from the faintly staining fundamental mass of the nucleus, aggregate gradually 
into a distinct number of larger masses, the chromocenters; (c) these chromo- 
centers unite to form larger masses, more compact and lengthened, which con- 
stitute the prochromosomes; (d) some prochromosomes fuse together by thet 
ends into a chain, which finally results in filaments of the spirem; the oak 
by this time is homogeneous in structure, but it is uncertain whether it is a single 
continuous thread; (e) the filaments of the spirem segment into a definite num 
ber of chromosomes, which seem to be more numerous than the prochromo- 
somes. (2) Metaphase: The chromosomes become arranged in the equatorial 
plate, where they divide and the daughter chromosomes separate. (3) Ana- 
se: The chromosomes accumulate in a convergent bundle at the pole. (4) 
Telophase: The chromosomes dissolve and form the fundamental nuclear mass 
with its chromatin granules.—Suicfo YAMANOUCHI. 
Riella.—In the twelfth of the Archegoniatienstudien, which he announces ' 
be the last, so far at least as concerns the liverworts, GOEBEL describes the Dr 
buds of Riella helicophylla, R. cossoniana, and R. Battandieri.17 These organs 
were first found by Unperwoop and Howe in R. americana, and have not a 
been observed in R. Clausonis. They consist of an unequally two-lobed dis 
attached by a single stalk cell somewhat excentrically placed on the upper eck 
face. GoErBEL holds these gemmae to be “‘ modified slime papillae,” as = 
chantia, an effort at homologizing which seems to us strained. The smaller lo 
1s loaded with food and therefore heavier; so the gemma sinks in the water 
with this end down and from it rhizoids arise. The larger lobe, which he calls 
sehen disk, grows, and especially the meristematic tissue between the 1 
parting them by a rather long stalk. From the germ disk two plants af at 
duced directly when well nourished, and otherwise one indirectly, that 35, : , 
fter proliferation of the germ disk. GorBet also discusses the systematic PO 
: F d 
tion of the genus, concluding on rather doubtful grounds, it seems, that it shoul 
: ovo 
? *© Baccarint, P., Sulle cinesi vegétative del Cynomorium coccineum TE 
Giorn. Bot. Ital. 15: 189-203. pl. 7. £268; ad 
: u 
*7 Gorsrt, K., Archegoniatenstudien. XII. Ueber die Brutknospenbilduné 
Uber die systematische Stellung von Riella. Flora 98;308-323- figs: 17: age? 
