a 
74 BOTANICAL GAZETTE [ FEBRUARY 
It is hardly necessary to refer to the history or literature oi 
this subject, as it has been presented frequently. The papet 
simply presents certain results obtained from Ranunculus, i 
reference to structures with whose “normal” behavior morpholo- 
gists are entirely familiar. It was felt that Ranunculus might be 
of interest from a phylogenetic point of view, as representing of 
of the so-called “ primitive regions” of dicotyledons. 
The usual methods of killing, staining, and sectioning wet 
used, and need no special description. 
MICROSPORANGIA AND MICROSPORES. g 
In the young sporangium a plate of hypodermal cells becomes 
distinctly differentiated by means of their enlarging radial diamett! 
and prominent nuclei (fig.7). There is great confusion in ter | 
minology in reference to these cells and those derived from them, 
and it is in the interest of clear statement to establish homolo 
gous parts, so far as possible, and adopt a uniform terminology: j 
In case this is not possible, at least a consistent terminology 
should be used throughout a single paper or text. Deferring® 
brief discussion of this matter to the end of the section, I woul!” 
regard this plate of cells as the archesporium. Each archespe 
rial cell tends to develop a radial row, and this arrangemelt 
remains more or less distinct in spite of certain inequalities ® 
a single archesporial cell, the sequence of events is vel 
lar to that commonly found in connection with the arches 
cell of the macrosporangium. A periclinal division result 
two cells radially placed (fig. r), the outer of which is a primal) | 
wall cell and the inner a primary sporogenous cell. From ti 
point, the sequence is variable. Both cells may divide by Fe 
clinal walls, or the primary sporogenous cell may divide 0" 7 
in the-formation of mother cells. In the former caseé, the out : 
aie formed by the periclinal division of the sporogenous cell cory 
peetes to the formation of the tapetum, In the latter case | 
innermost cell derived from the two or three periclinal divisio® 
of ths primary wall cell contributes to the tapetum (figs: ? ; 
