306 BOTANICAL GAZETTE [way 
1849, it was not until Hofmeister showed the independence of 
the rudimentary spore-producing organism from the thallus that 
a clear idea was possible of the early stages. Schacht’s fi. 6, 
jig. 2, of the work cited, shows a young sporophyte of about 
twenty cells, but drawn as if a branch of the thallus. The 
position of the walls, however, is correctly indicated. Hofmeis- 
ter, in 1851, cleared up the embryogeny of this plant so far as 
concerned the first segmentations. His i. z, fig. 36 of the work 
cited is practically correct, although not so much can be said for 
fig.39. It is to Leitgeb and Waldner that we owe the modern 
knowledge of the Anthoceros embryogeny. Leitgeb” first dis- 
cussed it fully in a special paper, and later Leitgeb and Wald- 
ner,” in their joint part of Leitgeb’s classic work on the Hepat- 
icze, described and figured with accuracy the young plants from 
the first segmentation on to maturity. In Leitgeb and Wald- 
ner’s memoir, f/. z, fig. z shows most clearly a young eight- 
celled stage, while earlier phases are de- 
scribed in the text. The originally formed 
wall is in this plant not the basal wall, 
but the median or transverse wall, run- 
ning parallel with the axis of - 08 
gone and dividing the egg into a 
halves, either anterior-posterior Of rig t 
and left, since the archegone pel 
Fic. 4.— Embryo of a perpendicular position bee ri 
Anthoceros. upper layers of the thallus. 1987 
the ovoid shape of the egg; sais ne 
plane of the first segmentation, and the later formed pee 
basal wall which divides the egg into capsular area and .@ imal 
quite as in the Marchantiacee. In this egg the ses ia in 
specialization is strongly marked. Not only does ae yee in 
the position of the basal wall below the middle of the ©88 " 
columnat 
the ovoid shape of the egg itself. The slender, 
Op. cit. 5. 1851. 
™ Entwickelung der Kapsel von Anthoceros. Sitz. der K. 
* Unters. iiber der Lebermoose 5:21. 
Acad. Wiss: 73 
