1898] THE LIFE HISTORY OF EUPHORLIA COROLLATA 419 
nally from a stamen only by a slight joint which marks the 
insertion of filament upon pedicel. In the center of the inflor- 
escence arises the pistillate flower, distinguished also from a pistil 
by a joint in its axis. It is made up of three carpels forming a 
trilocular ovary with a single suspended anatropous ovule in 
each chamber, and a style with three deeply bifid branches. 
Masses of glandular hairs envelop the base of. the pistillate 
flower, forming a dense screen between it and the staminate 
flowers. After fertilization the pedicel of the pistillate flower 
grows with great rapidity, carrying it far out of the involucre 
(fig. 12). 
ORGANOGENY. 
The first trace of the cyathium is a papilla-like outgrowth 
between two bracts which are themselves in the axil of a larger 
bract ( fig. z, and the bud shown in fig. 12). Near the base of 
this papilla there soon appear five protuberances (jig. 2) , the fore- 
tunners of the involucre. In their axils arise the oldest staminate 
flowers (fig. 3 5 t). Simultaneously with these, and shortly before 
the appearance of the carpels, the three ovules are distinguishable 
on the outer rim of the papilla (fg. 30). The three carpels 
develop independently from the main axis above the oldest 
‘Staminate flowers, which, at the same time, give off near their 
bases branches which are the second generation of staminate 
flowers (fig. 4). The carpels grow rapidly, their adjacent edges 
Soalescing and turning inward as they develop, in such a way as 
to enclose each of the three ovules in a separate compartment 
(Ag. 7). Athird set of staminate branches spring from the 
‘Pedicels of the second, and occasionally a fourth from these ( fig: 
The notch marking the boundary between filament and 
Pedice] appears when the stamen is nearly mature (jig. 5$*): 
A tegular perfect flower does not develop thus. Stamens 
‘Allse acropetally from the flower axis, and not as_ branches 
mM stamens of a preceding generation, as is evident aca 
from the course of the bundles. The resemblance of a single 
‘yathium to a reduced inflorescence of exactly the same type as 
