1899] THE COMPOUND OOSPHERE OF ALBUGO BLITI 163 
legniacee may be homologized with the central globule of the 
oosphere of Albugo. 
It is premature to discuss the function of this body until its 
relations in other species have been closely studied. As described 
by Wager it appears to be intimately connected with the behav- 
ior of the sexual nuclei; but there appears to be no such rela- 
tion in A. Buitt, where its function is perhaps that of an organizer 
in the oosphere. The body first appears when the oosphere is 
well defined and is most highly differentiated during the entrance 
of the daughter nuclei. At this time also occurs the formation of 
a thin film of denser protoplasm which definitely bounds the 
oosphere. A great dynamic change occurs in the oogonium 
When the ooplasm and periplasm are differentiated and the zone 
of cytoplasm separating the two regions is formed, and in addi- 
tion there is that remarkable division of the nuclei in such a 
manner that approximately fifty daughter nuclei are always cast 
into the ooplasm. Simultaneous with these activities, existing 
when they are at their maximum and disappearing when they 
®ease, there is formed this peculiar structure, which is so definite 
ne character and so constantly present that it seems to have some 
functional importance, 
In view of 
logical 
the wri 
the morphological character and possible physio- 
Value of this central structure to the coenocytic oosphere 
is bate . tures. to abel deus for it the name ce@nocentrum. It 
the iad ere to emphasize the difference in structure between 
Nucleus ea and a nucleus. The globule is distinctly not a 
Stains diffe it is much smaller than any nuclei that were seen, 
i » and is structureless and unchanging. The 
Since these " eer be a nucleus with the globule as a nucleolus, 
OF its [. fail to have the internal structure of a nucleus 
‘"8 membrane. There is no definite demarcation 
from ¢ : 
71,74, rounding ooplasm, as is plainly shown in figs. 47, 
In os 
— 75 other Structures are shown, the nature of which is 
uted in ik They are small granules which lie profusely distrib- 
“ cytoplasmic Strands in all ages of the oogonium, but 
he Sur 
75: 
. 
