168 BOTANICAL GAZETTE [SEPTEMBER 
to act conjointly with the normal elongating forces, thus pro- 
ducing extraordinarily long spindles (fig. 62). On the com 
trary, if the nuclei reach the border of the central mass ina 
earlier stage of mitosis no such forces obtain. Spindles caught 
in the first massing of the cytoplasm are often distorted ant 
bent like the letter 4 owing undoubtedly to torsion caused by 
the vacuoles as they move outward. ie 
MATURATION OF THE COMPOUND OOSPHERE AND OF THE 
ANTHERIDIUM. 
The multinucleate or compound oosphere when completely 
differentiated contains by actual count an average of 455 
nuclei. These are found in various conditions of reorganizatioa 
following the mitosis at zonation, and they rapidly assume th 
typical condition of a resting nucleus, each showing a promt 
nucleolus and very faint linin network. A nuclear membratteS 
sharply differentiated. Presently the linin network becom 
more prominent and a spirem condition is reached, ver Bi 
that first observed in the oogonium. A mitosis now one ae 
the oosphere affecting all of its nuclei, and is similar ba : 
important details to that just described for the oogonit 
as illustrated in jigs. 22-30. The nuclear figure stains 
more faintly than that of the previous division, the ; 
appearing lighter and skeleton-like in comparison with 
the first mitoses. The only other important differences 
“ests 
