1899] THE COMPOUND OOSPHERE OF ALBUGO BLITI 175 
time when an act of fertilization could possibly take place. 
Moreover, when only one or two nuclei are present they are 
always peripheral, which would not be expected if they resulted 
by the division of a fusion nucleus. Again, these scattered 
nuclei are always in the same condition as those near the 
outside of the developing oosphere, and this is almost invariably 
ametaphase of mitosis. This coincidence is inexplicable on the 
basis of their being the result of the division of a fusion nucleus, 
but it follows as a necessity from the explanation offered in this 
paper. If they are the product of one nucleus, which has under- 
gone three or four divisions, we have to assume not only the 
existence of the sex nuclei, their fusion, and the fusion nucleus, 
but also the resting, prophase, telophase, and anaphase conditions 
in the formation of 2, 4, 8, 16, and 32-celled stages. An anaphase 
nucleus is never seen in an oogonium except when the nuclei 
divide simultaneously either during the first or the second mitosis. 
The first anaphase always occurs when the general appearance 
S that shown in jig. 67,7. e., when the nuclei are completely 
lined upand the ooplasm is well differentiated. The second 
*pPeats in the oosphere after its complete differentiation ( figs. 70, 
4). It is impossible that the 50 nuclei of the oosphere can 
have been derived from a single hypothetical fusion nucleus. 
If attention is turned to the antheridial tube it might be sug- 
:. fertilization could take place at an early period, when 
ee eg are like those shown in figs. 62, 64. But it is only 
always ed ¥ plain the fact that the antheridial tube is 
tion cn: short at this time, and invariably occues the posi- 
Tarely if a. ae 64. The wall of the tube is thick, the oe 
Cation that m contains nuclei, and there is not the slightest indi- 
that it ae y at all ready toopen. If it be assumed, however, 
Nucleus that release, in some manner difficult to detect, a single 
female eke, ad the male nucleus, and which fuses with we 
e develo is Ow can the continued growth of the tube an 
70,73 i of such peculiar conditions as are shown in ~~ 
tube a” 50, 82, 83, etc., be explained? Why does the 
grow after functioning only to meet the difficulty 
