1899] THE COMPOUND OOSPHERE OF ALBUGO BLITI 237 
in earlier pages the broader significance of the phenomena 
described. 
A mitotic division of the nuclei of fungi seems to have been 
first noticed by Sadeback (’83) in Exoascus, and has since been 
observed in various forms and carefully studied and described in 
_ afew papers. 
The occurrence of a mitotic division in the oogonium of 
_ Albugo was discovered by Wager (96) in A. candidus, and con- 
firmed by Berlese (98) in A. Portulace. Wager notes that the 
muclei enlarge and divide, leaving one daughter nucleus imbedded 
inthe central body while the other nuclei pass to the periplasm. 
The one division increases the nuclei from about 115 to about 
double that number. Berlese says the nuclei divide several 
times in A. Portulace, increasing the number from 30 or 40 to 
about 200. The account given in the present paper describes two 
*pproximately simultaneous divisions affecting all of the nuclei 
® both antheridium and oogonium, and these mitoses result in 
ie formation of the sexual elements, numerous male and female 
nuclei, The second division is strikingly different from the first 
Be erence of the nuclear elements, particularly the 
tna of = This condition suggested the possibility of a sgeieee 
evidence a. but careful study revealed na: Conve se 
a ee - The mitoses are characterized by the intranuclear 
| ation of the spindle, the intranuclear centrosomes, the per- 
5 ~Maence of the nucleolus, and the entire absence of extra- 
 ‘Welear radiation. 
get described the disappearance of the nucleolus in early 
followed by the formation of chromosomes and then 
- the spin B  vclopment. He inclines toward the view that 
WW to bia ‘ €rived from the linin. The membrane persists 
ie ‘iad 2 and Wager did not follow the division seem 
a dissppea, Observations similar to those of Wager, pein 
indle a of the nucleolus in prophase, argues that ; : 
in 4. Blin tf der ived from it. The behavick of the nuc eo 
"entioneg ; 's different from that described by the writers 
: » in that the structure remains apparent inside of the 
a aia i ial 
